- Single-point tubercle, rarely absent, at each funnel-mantle fusion.
- Arms II of males with 2 series of suckers on modified tips (sometimes appearing as 3-4 series due to crowding).
- Arms II of males with 15-19 pairs of normal suckers proximal to modified tips.
- Arms I of males without modified tips.
- Largest arm III suckers 3 times basal suckers in diameter (at least 2.0-2.8% of GL in diameter).
- Largest club suckers with 19-24 teeth.
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