Thallus: foliose, orbicular to irregularly spreading, 3-7(-9) cm in diam, loosely attached centrally, with free and sub-ascendent margins, rather coriaceous; lobes: irregularly laciniate, rather ragged, 10-20(-30) mm wide, 10-30(-35) mm long, discrete to subimbricate at margins, complex-imbricate centrally; margins: entire, sinuous, here and there, shallowly notched or scalloped, thickened below with a distinct, dark rim; upper surface: when wet dark lead-gray to blue-black marginally and paler centrally, when dry pale gray-blue or glaucous gray and suffused brown, with minute, white, marbled maculae (x10 lens, moist), smooth or shallowly wrinkled, often with minute, pale-buff or white, scattered to crowded papillae (x10 lens), dull, without isidia, phyllidia or soredia; medulla: white, K-; photobiont: cyanobacterial; lower surface: pale-buff to chocolate brown, tomentose from margins to center, sparse and rather cobwebby, to entangled-woolly, rhizinate; rhizines: occasionally with fascicles of long rhizines (up to 4 mm long) centrally, also here and there with projecting, solitary to fasciculate brown-black rhizines up to 1 mm long; cyphellae: common, scattered, prominent, minute, pin-prick-like at margins, round to ellipsoid, (0.1-)0.3-1(-1.5) mm in diam. centrally, with flat, pale, "halo-like" margins, with a white basal membrane; Apothecia: occasional to frequent, marginal and laminal, best developed and often clustered towards lobe apices, sessile, constricted at base, round, (0.5-)1-2(-2.5) mm in diam.; disc: orange to orange-brown and translucent when moist, red-brown and dull when dry, epruinose; margins: pale pinkish buff to cream-colored, coarsely corrugate, scabrid, obscuring disc at first, persisting, crenulate to ragged at maturity, often with projecting, silky, glistening, white to brownish cilia 0.2-0.6(-0.8) mm long); epihymenium: pale yellow-brown to orange-brown, 7-12 µm thick; hymenium: hyaline, 75-95 µm tall; hypothecium; opaque, yellow-brown, 50-80 µm thick; asci: clavate, 55-70 x 10-13.5 µm, 8-spored; ascospores: hyaline to pale yellow-brown, 1-3-septate, fusiform-ellipsoid, with pointed apices, 25-30(35) x 5.5-8.5(-10) µm; Pycnidia: occasional, scattered, developed towards margins, immersed but apparent as distinct, pale buff or pale brown swellings, 0.2-0.5(-0.8) mm in diam., with a central, depressed red-brown to brown-black ostiole; conidia: bacilliform, straight, 3-5 x 1 µm; Spot tests: all negative; Secondary metabolites: none detected.; Substrate and ecology: pine-oak forest, on Quercus, 2320 m; World distribution: neotropical in Mexico, Colombia, Venezuela, Peru, Brazil; Sonoran distribution: Sinaloa (only one collection seen).
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