Non-Migrant: No. All populations of this species make significant seasonal migrations.
Locally Migrant: Yes. At least some populations of this species make local extended movements (generally less than 200 km) at particular times of the year (e.g., to breeding or wintering grounds, to hibernation sites).
Locally Migrant: Yes. At least some populations of this species make annual migrations of over 200 km.
Anadromous. Adults return to spawn in areas where they were hatched. Individuals may move up to 2,000 km upstream to spawn in rivers lacking major barriers (Lee et al. 1980).