Shell large, rather thick and solid. Umbones subterminal, at about one-tenth of the shell length. Ligament strong. Extremely thick and large ctenidia with long demibranchs. No extreme mantle fusion, valvular siphonal membrane short. Anterior adductor muscle scar located in front of the umbo. Posterior adductor muscle scar rounded trapezoid. Gut with a single clockwise loop.
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