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 Thallus: foliose, orbicular to irregular, fragile, up to 2 cm diam, ± firmly adnate, central parts often covered by pustules dissolved into granulate soredia; lobes: flat to concave, up to 0.5 mm broad, rarely broader, sparsely divided; tips: truncate or crenulate, occasionally ascending; upper surface: whitish, rarely dark gray, usually dull; pustules or isidia: laminal, often developing into granular soralia or craters that may fuse and cover the whole thallus; upper cortex: paraplectenchymato; lower cortex: ± prosoplectenchymatous, pale to brownish (inner parts); lower surface: white at the tips, darker in inner parts; rhizines: few, concolorous with the underside; Apothecia: frequent, up to 2 mm diam., margin often crenulate; ascospores: brown, 1-septate, Pachysporaria type, (16-) 18.5-22 (-23.5) x (7.5-) 8.5-10 (-11) µm; Pycnidia: not seen; Spot tests: upper cortex and medulla K+ yellow, C-, KC-, P+ yellow; Secondary metabolites: upper cortex with atranorin; medulla with atranorin, lecotylin and traces of other triterpenoids visible in TLC.; Substrate and ecology: growing mainly on trees but also on rocks in open sites; World distribution: known from scattered localities in central and southern Europe and from southern California; Sonoran distribution: known only from the Channel Islands of southern California and the adjacent mainland. 


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© Lichen Unlimited: Arizona State University, Tempe.

Source: Lichen Flora of the Greater Sonoran Desert Region

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