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 Thallus: adnate, foliose, 4-20 cm in diam., lobate; lobes: sublinear, contiguous to imbricate, elongate, plane, separate, 2-4 mm wide, apices truncate; upper surface: gray, smooth to strongly foveolate, shiny, sometimes white pruinose, becoming cracked along prominent, effigurate pseudocyphellae; soredia: granular, abundant, laminal, in linear soralia; isidia, pustulae and dactyls absent; medulla: white with continuous algal layer; lower surface: black, rhizinate; rhizines moderately dense, black, simple; Apothecia: rare, laminal, 2-8 mm wide; margin: usually sorediate; disc: dark brown; asci: clavate, 8-spored; ascospores: ellipsoid, 11-14 x 6-8 µm; Pycnidia: infrequent, immersed; conidia: bacilliform, 6-8 x 1 µm; Spot tests: upper cortex K+ yellow, C-, KC-, P+ yellow; medulla K+ yellow turning deep red, C-, KC-, P+ orange; Secondary metabolites: upper cortex with atranorin and chlorotranorin; medulla with salazinic acid (major) and consalazinic acids (minor).; Substrate and ecology: common on trees in open habitats, occasionally on rocks; World distribution: pantemperate and southern boreal; Sonoran distribution: relatively common on deciduous and coniferous trees in montane habitats of Arizona, on coastal oaks in California and on rocks in the high mountains of northern Baja California. 


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© Lichen Unlimited: Arizona State University, Tempe.

Source: Lichen Flora of the Greater Sonoran Desert Region

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