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Range DescriptionThis species ranges from the eastern side of the Balkan Peninsula (extreme southeastern Serbia, eastern FYR Macedonia, most of Bulgaria, northeastern Greece, and European Turkey) through western and northern Turkey along the edges of the Black Sea, to the Caucasus (Russia, Georgia and Azerbaijan). There are apparently isolated populations in central Serbia, on the Crimean Peninsula (Ukraine), and in the southern Caspian region of northern Iran. The map for this species largely follows Arntzen, (2003). It can be found from sea level to elevations approaching 2,100m asl (southern Caucasus).