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Holotype: USNM 7347 according to Golay et al., 1993, Endogly. Venom. Snakes World 478pp.[177].

Type-locality: Taboga Island, Bay of Panama.

Many subspecies have been described; most of these are ill-defined with a considerable amount of overlap of diagnostic characters occurring between populations. The following subspecies were recognized by Roze, 1983 [dated 1982], Mem. Inst. Butantan 46:305-338: M. n. nigrocinctus (Girard, 1854); M. n. babaspul Roze, 1967; M. n. coibensis Schmidt, 1936; M. n. divaricatus (Hallowell, 1855); M. n. melanocephalus (Hallowell, 1860); M. n. mosquitensis Schmidt, 1933; M. n. ruatanus (Günther, 1895), and M. n. zunilensis Schmidt, 1932.

M. ruatanus was recognized as a valid species by Wilson and Meyer, 1985, Snakes Honduras, 2nd ed., 159pp.[115]; Campbell and Lamar, 1989, Venom Rept. Latin Am., 425pp.[142]; and Wilson, McCranie, and Slowinski, 1992, Cat. Am. Amph. Rept. 545. 1-2[1].

There are probably several species in this complex; several populations were shown to have chromosomal differences by Gutérrez and Bolaños, 1979, Rev. Biol. Trop. 27:57-73, and Gutiérrez and Bolaños, 1981 [dated 1980], Rev. Biol. Trop. (1)29:115-122.

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© Mohammadi, Shabnam

Source: Snake Species of the World

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