James E. Lotan and William B. Critchfield
Lodgepole pine (Pinus contorta) is a two-needled pine of the subgenus Pinus. The species has been divided geographically into four varieties: P. contorta var. contorta, the coastal form known as shore pine, coast pine, or beach pine; P. contorta var. bolanderi, a Mendocino County White Plains form in California called Bolander pine; P. contorta var. murrayana in the Sierra Nevada, called Sierra lodgepole pine or tamarack pine; and P. contorta var. latifolia, the inland form often referred to as Rocky Mountain lodgepole pine or black pine. Although the coastal form grows mainly between sea level and 610 m (2,000 ft), the inland form is found from 490 to 3660 m (1,600 to 12,000 ft).
Global Range: Extensive range from Alaska to northern New Mexico and Baja California, attaining best populations in the Rocky Mountains (Record and Hess 1943).
- The native range of lodgepole pine.
Puget Lowland Forests Habitat
Cope's giant salamander is found in the Puget lowland forests along with several other western North America ecoregions. The Puget lowland forests occupy a north-south topographic depression between the Olympic Peninsula and western slopes of the Cascade Mountains, extending from north of the Canadian border to the lower Columbia River along the Oregon border. The portion of this forest ecoregion within British Columbia includes the Fraser Valley lowlands, the coastal lowlands locally known as the Sunshine Coast and several of the Gulf Islands. This ecoregion is within the Nearctic Realm and classified as part of the Temperate Coniferous Forests biome.
The Puget lowland forests have a Mediterranean-like climate, with warm, dry summers, and mild wet winters. The mean annual temperature is 9°C, the mean summer temperature is 15°C, and the mean winter temperature is 3.5°C. Annual precipitation averages 800 to 900 millimeters (mm) but may be as great as 1530 mm. Only a small percentage of this precipitation falls as snow. However, annual rainfall on the San Juan Islands can be as low as 460 mm, due to rain-shadow effects caused by the Olympic Mountains. This local rain shadow effect results in some of the driest sites encountered in the region. Varied topography on these hilly islands results in a diverse assemblage of plant communities arranged along orographically defiined moisture gradients. Open grasslands with widely scattered trees dominate the exposed southern aspects of the islands, while moister dense forests occur on northern sheltered slopes characterized by Western red cedar (Thuja plicata), Grand fir (Abies grandis), and Sword fern (Polystichum munitum) communities.
There are only a small number of amphibian taxa in the Puget lowland forests, namely: Cope's giant salamander (Dicamptodon copei); Monterey ensatina (Ensatina eschscholtzii); Long-toed salamander (Ambystoma macrodactylum); Western redback salamander (Plethodon vehiculum); Northwestern salamander (Ambystoma gracile); Pacific chorus frog (Pseudacris regilla); Coastal giant salamander (Dicamptodon tenebrosus); Rough-skin newt (Taricha granulosa); the Vulnerable Spotted frog (Rana pretiosa); Tailed frog (Ascopus truei); and Northern red-legged frog (Rana aurora).
Likewise there are a small number of reptilian taxa within the ecoregion: Common garter snake (Thamnophis sirtalis); Western terrestrial garter snake (Thamnophis sirtalis); Northern alligator lizard (Elgaria coerulea); Western fence lizard (Sceloporus occidentalis); Northwestern garter snake (Thamnophis ordinoides); Sharp-tailed snake (Contia tenuis); Yellow-bellied racer (Coluber constrictor); and Western pond turtle (Clemmys marmorata).
There are numberous mammalian taxa present in the Puget lowland forests. A small sample of these are:Creeping vole (Microtus oregoni), Raccoon (Procyon lotor), Southern sea otter (Enhydra lutris), Mink (Mustela vison), Coyote (Canis latrans), Black-tailed deer (Odocoileus hemionus), Pallid bat (Antrozous pallidus), and Harbour seal (Phoca vitulina).
A rich assortment of bird species present in this ecoregion, including the Near Threatened Spotted owl (Strix occidentalis), Turkey vulture (Cathartes aura), Bald eagle (Haliaeetus leucocephalus), Blue grouse (Dendragapus obscurus), as well as a gamut of seabirds, numerous shorebirds and waterfowl.
Sierra Nevada Forests
The Limestone salamander is a highly localized endemic of the Sierra Nevada forests foothills conifned to a limited reach of the Merced River. The Sierra Nevada forests are the forested areas of the Sierra Nevada Mountains, which run northwest to southwest and are approximately 650 kilometers long and 80 km wide. The range achieves its greatest height towards the south, with a number of peaks reaching heights of over 4000 meters. Several large river valleys dissect the western slope with dramatic canyons. The eastern escarpment is much steeper than the western slope, in general.
The Sierra Nevada forests ecoregion harbors one of the most diverse temperate conifer forests on Earth displaying an extraordinary range of habitat types and supporting many unusual species. Fifty percent of California's estimated 7000 species of vascular plants occur in the Sierra Nevada, with 400 Sierra endemics and 200 rare species. The southern section has the highest concentration of species and rare and endemic species, but pockets of rare plants occur throughout the range.
Sierra Nevada amphibian endemics are the Yosemite toad, Mount Lyell salamander (Hydromantes platycephalus), the Vulnerable Limestone salamander (Hydromantes brunus), Kern salamander and the Endangered Inyo Mountains salamander (Batrachoseps campi). The non endemic amphibians are: the Endangered Southern mountain yellow-legged frog (Rana muscosa); the Near Threatened Cascades frog (Rana cascadae); Northern red-legged frog (Rana aurora); Pacific chorus frog (Pseudacris regilia); Foothill Yellow-legged frog (Rana boylii); Long-toed salamander (Ambystoma macrodactylum); and the Monterey ensatina (Ensatina eschscholtzii).
A considerable number of mammalian taxa are found in the ecoregion, including the Long-eared chipmunk, Alpine chipmunk, Western heather vole, Walker Pass pocket mouse, and the Yellow-eared pocket-mouse. A diverse vertebrate predator assemblage once occurred in the ecoregion including Grizzly bear (Ursus arctos), Black bear (Ursus americanus), Coyote (Canis latrans), Mountain lion (Puma concolor), Ringtail (Bassariscus astutus), Fisher (Martes pennanti), Pine marten (Martes americana) and Wolverine (Gulo gulo).
There are a small number of reptilian taxa present in the Sierra Nevada forests: sagebrush lizard (Sceloporus graciosus); Northern alligator lizard (Elgaria coerulea); Southern alligator lizard (Elgaria multicarinata); Sharp-tailed snake (Contia tenuis); California mountain kingsnake (Molothrus ater); Common garter snake (Thamnophis sirtalis); Couch's garter snake (Thamnophis couchii); Western gopher snake (Pituophis catenifer); Longnose snake (Rhinocheilus lecontei); and the Common kingsnake (Lampropeltis getula).
A number of bird species are found in the ecoreion including high level predators that include several large owls, hawks and eagles. Other representative avifauna species present are the Blue-headed vireo (Vireo solitarius); Brown-headed cowbird (Molothrus ater); and the Near Threatened Cassin's finch (Carpodacus cassinii).
Sierra Juarez and San Pedro Martir pine-oak forests Habitat
This taxon can be found in the Sierra Juarez and San Pedro Martir pine-oak forests. The ecoregion is located in two mountain ranges in the state of Baja California, Mexico: the Sierra de Juarez and the Sierra de San Pedro Martir. Both mountain ranges belong to the physiographical province of Baja California, and constitute the northernmost elevated peaks of the Baja Peninsula. The mountainous range that descends into a large portion of Baja California becomes more abrupt at Juarez and San Pedro Martir; the eastern slope is steeper than the western. Altitudes range between 1100-2800 meters. The granitic mountains of Juarez and San Pedro Martir have young rocky soils and are poorly developed, shallow, and low in organic matter.
Dominant trees in the ecoregion are: Pinus quadrifolia, P. jeffreyi, P. contorta, P. lambertiana, Abies concolor, and Libocedrus decurren. The herbaceous stratum is formed by Bromus sp. and Artemisia tridentata. Epiphytes and fungi are abundant throughout the forests.
Characteristic mammals of the ecoregion include: Ornate shrew (Sorex ornatus), Puma (Puma concolor), Fringed Myotis bat (Myotis thysanodes), California chipmunk (Tamias obscurus), Bobcat (Lynx rufus), Coyote (Canis latrans), San Joaquin kit fox (Vulpes macrotis) and Bighorn sheep (Ovis canadensis).
Numerous birds are present in the ecoregion, including the rare Bald eagle (Haliaeetus leucocephalus), California condor (Gymnogyps californianus), Pinyon jay (Gymnohinus cyanocephalus), and White-breasted nuthatch (Sitta carolinensis).
A number of different reptilian taxa are found in these oak-pine forests; representative reptiles here are: the Banded rock lizard (Petrosaurus mearnsi); Common checkered whiptail (Cnemidophorus tesselatus), who is found in sparsely vegetated areas; Coast horned lizard (Phrynosoma coronatum), often found in locales of sandy soil, where individuals may burrow to escape surface heat; Night desert lizard (Xantusia vigilis), who is often found among bases of yucca, agaves and cacti; and the Baja California spiny lizard (Sceloporus zosteromus).
The Pacific chorus frog (Pseudacris regilla) is an anuran found within the Sierra Juarez and San Pedro Martir pine-oak forests as one of its western North America ecoregions of occurrence. The only other amphibian in the ecoregion is the Western toad (Anaxyrus boreas).
California Montane Chaparral and Woodlands Habitat
This taxon can be found in the California montane chaparral and woodlands, a near coastal ecoregion in Central and Southern California, USA. This ecoregion is disjunctive, with a major element in Southern California and another along the Monterey County coast. The ecoregion encompasses most of the Transverse Range that includes the San Bernardino Mountains; San Gabriel Mountains; portions of the Santa Ynez and San Rafael Mountains; Topatopa Mountains; San Jacinto Mountains; the Tehachapi, Greenhorn, Piute, and Kiavah Mountains that extend roughly northeast-southwest from the southern Sierra Nevada; and the Santa Lucia Range that parallels the coast southward from Monterey Bay to Morro Bay.
The California montane chaparral and woodland ecoregion consists of a complex mosaic of coastal sage scrub, lower chaparral dominated by chamise, upper chaparral dominated by manzanita, desert chaparral, Piñon-juniper woodland, oak woodlands, closed-cone pine forests, yellow pine forests, sugar pine-white fir forests, lodgepole pine forests, and alpine habitats. The prevalence of drought-adapted scrub species in the flora of this ecoregion helps distinguish it from similar communities in the Sierras and other portions of northern California. Many of the shared Sierra Nevadan species typically are adapted to drier habitats in that ecoregion, Jeffrey Pine (Pinus jeffreyi) being a good example.
Oak species are an important component of many chaparral and forest communities throughout the ecoregion. Canyon Live Oak, Interior Live Oak, Tanbark Oak (not a true Quercus species), Engelmann Oak, Golden-cup Oak, and Scrub Oak are some examples. Mixed-conifer forests are found between 1371 to 2896 meters elevation with various combinations and dominance of incense cedar, sugar pine, and white fir, Jeffrey Pine, Ponderosa Pine, and mountain juniper. Subalpine forests consist of groves of Limber Pine (Pinus flexilis), Lodgepole Pine, and Jeffrey Pine. Very old individual trees are commonly observed in these relict subalpine forests. Within this zone are subalpine wet meadows, talus slope herbaceous communities, krumholz woodlands, and a few small aspen groves.
In addition to these general vegetation patterns, this ecoregion is noted for a variety of ecologic islands, communities with specialized conditions that are widely scattered and isolated and typically harbor endemic and relict species. Examples include two localities of Knobcone Pine (Pinus attenuata) on serpentine soils, scattered vernal pools with a number of endemic and relict species, and isolated populations of one of North America’s most diverse cypress floras, including the rare Gowen Cypress (Cupressus goveniana goveniana) restricted to two sites on acidic soils in the northern Santa Lucia Range, Monterey Cypress (Cupressus macrocarpa) found only at two coastal localities near Monterey Bay, and Sargent Cypress (Callitropsis sargentii LR/LC) restricted to serpentine outcrops. Monterey Pine (Pinus radiata) is also restricted to three coastal sites near Monterey Bay.
The ecoregion is also home to a few endemic or near-endemic mammalian vertebrates, such as the White-eared Pocket Mouse (Perognathus alticolus EN), a mammal known only to two disjunct mountain ranges in southern California: San Bernardino Mountains in San Bernardino County (ssp. alticolus), and the Tehachapi Mountains, in Kern, Ventura, and Los Angeles counties. The near-endemic fossorial Agile Kangaroo Rat (Dipodomys agilis) is found in the southern disjunctive unit of the ecoregion, and is known only to the Los Angeles Basin and foothills of San Gabriel and San Bernardino mountains in Ventura, Los Angeles, and Riverside counties north to Santa Barbara County and through the southern Sierra Nevada, including Mount Pinos, Tehachapi and San Gabriel mountains, and northern San Fernando Valley. Non-endemic mammals found in the ecoregion include Botta's Pocket Gopher (Thomomys bottae) and Trowbridge's Shrew (Sorex trowbridgii). Some larger vertebrate predators can be found in the ecoregion, including Puma (Puma concolor), Bobcat (Lynx rufus), Coyote (Canis latrans), and Ringtails (Bassariscus astutus).
The ecoregion boasts five endemic and near-endemic amphibians, largely Plethodontid salamanders. Some specific salamander taxa found here are the endemic Tehachapi Slender Salamander (Batrachoseps stebbinsi VU), known from isolated sites in the Caliente Creek drainage, Piute Mountains, and Kern County, California along with scattered populations in the Tehachapi Mountains to Fort Tejon, Kern County; the near-endemic Blackbelly Slender Salamander (Batrachoseps nigriventris); the Monterey Ensatina (Ensatina eschscholtzii); the Channel Islands Slender Salamander (Batrachoseps pacificus), endemic to a narrow range restricted solely on Anacapa, Santa Cruz, Santa Rosa, and San Miguel islands; and the Arboreal Salamander (Aneides lugubris), found only in California and Baja California. A newt found here is the Coast Range Newt (Taricha torosa). Anuran taxa in the ecoregion include the Foothill Yellow-legged Frog (Rana boylii NT); the Southern Mountain Yellow-legged Frog (Rana muscosa EN), a California endemic occurring in several disjunctive populations; and the Northern Red-legged Frog (Rana aurora).
The California montane chaparral and woodlands ecoregions contains a number of reptiles such as the Coast Horned Lizard (Phrynosoma coronatum), who ranges from Northern California to Baja California. Also found here is the Sagebrush Lizard (Sceloporus graciosus); the Western Fence Lizard (Sceloporus occidentalis); the Southern Alligator Lizard (Elgaria multicarinata); and the Side-blotched Lizard (Uta stansburiana). The Two-striped Garter Snake (Thamnophis hammondii) is a restricted range reptile found near-coastally from Monterey County, California southward to Baja California.
The California Condor once inhabited much of the ecoregion, with the western Transverse Range acting today as a refuge for some of the last wild populations, after considerable conservation efforts, especially in the Los Padres National Forest. The Heermann's Gull (Larus heermanni NT) is found in coastal areas of the ecoregion.
Arizona Mountains Forests Habitat
This taxon is found in the Arizona Mountain Forests, which extend from the Kaibab Plateau in northern Arizona to south of the Mogollon Plateau into portions of southwestern Mexico and eastern Arizona, USA. The species richness in this ecoregion is moderate, with vertebrate taxa numbering 375 species. The topography consists chiefly of steep foothills and mountains, but includes some deeply dissected high plateaus. Soil types have not been well defined; however, most soils are entisols, with alfisols and inceptisols in upland areas. Stony terrain and rock outcrops occupy large areas on the mountains and foothills.
The Transition Zone in this region (1980 to 2440 m in elevation) comprises a strong Mexican fasciation, including Chihuahua Pine (Pinus leiophylla) and Apache Pine (P. engelmannii) and unique varieties of Ponderosa Pine (P. ponderosa var. arizonica). Such forests are open and park-like and contain many bird species from Mexico seldom seen in the U.S.. The Canadian Zone (above 2000 m) includes mostly Rocky Mountain species of mixed-conifer communities such as Douglas-fir (Pseudotsuga menziesii), Engelmann Spruce (Picea engelmanni), Subalpine Fir (Abies lasiocarpa), and Corkbark Fir (A. lasiocarpa var. arizonica). Dwarf Juniper (Juniperus communis) is an understory shrubby closely associated with spruce/fir forests. Exposed sites include Chihuahua White Pine (Pinus strobiformis), while disturbed north-facing sites consists primarily of Lodgepole Pine (Pinus contorta) or Quaking Aspen (Populus tremuloides).
There are a variety of mammalian species found in this ecoregion, including the endemic Arizona Gray Squirrel (Sciurus arizonensis), an herbivore who feeds on a wide spectrum of berries, bark and other vegetable material. Non-endemic mammals occurring in the ecoregion include: the Banner-tailed Kangaroo Rat (Dipodomys spectabilis NT); Desert Pocket Gopher (Geomys arenarius NT). In addition, there is great potential for restoring Mexican Wolf (Canis lupus) and Grizzly Bear (Ursus arctos horribilis) populations in the area because of its remoteness and juxtaposition to other ecoregions where these species were formerly prevalent.
There are few amphibians found in the Arizona mountain forests. Anuran species occurring here are: Red-spotted Toad (Anaxyrus punctatus); Southwestern Toad (Anaxyrus microscaphus); New Mexico Spadefoot Toad (Spea multiplicata); Woodhouse's Toad (Anaxyrus woodhousii); Northern Leopard Frog (Lithobates pipiens); Chiricahua Leopard Frog (Lithobates chiricahuensis VU); Madrean Treefrog (Hyla eximia), a montane anuran found at the northern limit of its range in this ecoregion; Boreal Chorus Frog (Anaxyrus woodhousii); Western Chorus Frog (Pseudacris triseriata); and Canyon Treefrog (Hyla arenicolor). The Jemez Mountains Salamander (Plethodon neomexicanus NT) is an ecoregion endemic, found only in the Jemez Mountains of Los Alamos and Sandoval counties, New Mexico. Another salamander occurring in the ecoregion is the Tiger Salamander (Ambystoma tigrinum).
A number of reptilian taxa occur in the Arizona mountains forests, including: Gila Monster (Heloderma suspectum NT), often associated with cacti or desert scrub type vegetation; Narrow-headed Garter Snake (Thamnophis rufipunctatus), a near-endemic found chiefly in the Mogollon Rim area; Sonoran Mud Turtle (Kinosternon sonoriense NT).
Habitat and Ecology
Comments: Thrives in mostly well drained soils but may be found in peat bogs, muskegs or on dry sandy sites (Tree Talk 1994).
Soils and Topography
Extensive stands of lodgepole pine (var. latifolia) occur on soils classified as Inceptisols or Alfisols in the interior forests. Although the species commonly grows on Andepts and does well on these soils in some areas, the Boralfs and Ochrepts probably support better tree development and more extensive stands. Frequently lodgepole pine soils on Boralfs and Ochrepts have cryic soil temperature regimes. In the Blue Mountains of Oregon lodgepole pine does well on Andepts, where it is nearly always found on volcanic ash or alluvial material overlying residual basaltic soils, at elevations between 910 and 2130 m (3,000 and 7,000 ft). The ash cap soils are deeper and hold more moisture than the residual soils.
The coastal form of lodgepole pine (var. contorta) is often found on Histosols (peat bogs or muskegs) in southeastern Alaska, British Columbia, and western Washington, and on dry, sandy, or gravelly sites farther south along the coast on Inceptisols, Alfisols, and Ultisols.
Soil properties and soil moisture often favor lodgepole pine locally over other species. Lodgepole pine grows on wet flats and poorly drained soils in the Cascade Range in Washington and Oregon, and the Sierra Nevada in California. Soils with underlying hardpan support lodgepole pine to the exclusion of such species as ponderosa pine (Pinus ponderosa), redwood (Sequoia sempervirens), or Douglas-fir in the Sierra Nevada, eastern Oregon, and Mendocino County, CA. Lodgepole pine also grows on level sites with and without high water tables in central Oregon where frost tolerance during germination allows its establishment to the exclusion of other species. Extensive stands are found in these areas on well drained sites above 1600 m (5,250 ft), with patterns of occurrence attributed to past fires.
On infertile soils, lodgepole pine is often the only tree species that will grow. Nevertheless, experiments have demonstrated significant growth increase from fertilization, particularly nitrogen (15).
Lodgepole pine thrives in a wide variety of topographic situations. It grows well on gentle slopes and in basins, but good stands are also found on rough and rocky terrain and on steep slopes and ridges, including bare gravel. Northern and eastern slopes are more favorable than southern and western aspects (3).
At low elevations in the interior, lodgepole pine grows in areas receiving only 250 mm (10 in) of mean annual precipitation, whereas it receives more than 500 mm (200 in) along the northern coast. Many interior sites often are low in summer rainfall. Seasonal distribution of precipitation is significant; snowfall supplies most of the soil water used for rapid growth in early summer. Temperatures are frequently favorable for germination after snowmelt, and germination occurs rapidly. Lodgepole is very intolerant of shade and generally grows best in full sunlight.
aecium of Coleosporium asterum parasitises live Pinus contorta
In Great Britain and/or Ireland:
Foodplant / mycorrhiza / ectomycorrhiza
fruitbody of Cortinarius malicorius is ectomycorrhizal with live root of Pinus contorta
Remarks: Other: uncertain
Other: major host/prey
Foodplant / saprobe
fruitbody of Dacryobolus sudans is saprobic on decayed wood of Pinus contorta
Foodplant / pathogen
Brunchorstia anamorph of Gremmeniella abietina infects and damages live twig of Pinus contorta
Other: unusual host/prey
Plant / associate
fruitbody of Hebeloma arenosa is associated with Pinus contorta
Foodplant / internal feeder
larva of Hylobius abietis feeds within dead stump of Pinus contorta
Other: major host/prey
Foodplant / sap sucker
nymph of Leptoglossus occidentalis sucks sap of unripe seed (in 1-year old cone) of Pinus contorta
Remarks: season: 5-8
Other: major host/prey
Foodplant / saprobe
Cryptosporiopsis anamorph of Pezicula livida is saprobic on dead, fallen branch of Pinus contorta
Foodplant / pathogen
fruitbody of Phaeolus schweinitzii infects and damages live root of mature tree of Pinus contorta
Other: minor host/prey
Foodplant / pathogen
pycndium of Ramichloridium anamorph of Ramichloridium pini infects and damages shoot (young) of Pinus contorta
Fungus / saprobe
conidioma of Sirococcus coelomycetous anamorph of Sirococcus conigenus is saprobic on fallen cone of Pinus contorta
Foodplant / saprobe
effuse colony of Troposporella dematiaceous anamorph of Troposporella monospora is saprobic on dead needle of Pinus contorta
Associated Forest Cover
Lodgepole pine is a component in 27 of the 55 SAF western forest cover types. In the Northern Interior (Boreal) group it is represented in White Spruce (Type 201), White Spruce-Aspen (Type 251), White Spruce-Paper Birch (Type 202), Paper Birch (Type 252), and Black Spruce (Type 204).
It is a component in all six high elevation cover types: Mountain Hemlock (Type 205), Engelmann Spruce-Subalpine Fir (Type 206), Red Fir (Type 207), Whitebark Pine (Type 208), Bristlecone Pine (Type 209), and California Mixed Subalpine (Type 256). At middle elevations in the interior it is a minor component of seven other types: Interior Douglas-Fir (Type 210), Western Larch (Type 212), Grand Fir (Type 213), Western White Pine (Type 215), Blue Spruce (Type 216), Aspen (Type 217), and Limber Pine (Type 219). In the North Pacific forests, it is a component in Coastal True Fir (Type 226), Western Redcedar-Western Hemlock (Type 227), Western Redcedar (Type 228), Douglas-Fir-Western Hemlock (Type 230), Port-Orford-Cedar (Type 231), and Redwood (Type 232). At low elevations in the interior it is associated with Interior Ponderosa Pine (Type 237) and in the South Pacific forests it is a component of Jeffrey Pine (Type 247).
Lodgepole pine, with probably the widest range of environmental tolerance of any conifer in North America, grows in association with many plant species (30,50,59,60). The lodgepole pine forest type is the third most extensive commercial forest type in the Rocky Mountains.
Lodgepole pine's successional role depends upon environmental conditions and extent of competition from associated species. Lodgepole pine is a minor seral species in warm, moist habitats and a dominant seral species in cool dry habitats. It is often persistent even on cool and dry sites and can attain edaphic climax at relatively high elevations on poor sites. Fire regimes have played a role in this successional continuum, especially where repeated fires have eliminated a seed source for other species (27). Lodgepole pine may even overwhelm a site with seed stored in serotinous cones. It has four basic successional roles (50):
Minor Seral- A component of even-aged stands rapidly being replaced by shade-tolerant associates in 50 to 200 years.
Dominant Seral- The dominant cover type of even-aged stands with a vigorous understory of shade-tolerant species that will replace lodgepole pine in 100 to 200 years.
Persistent- The dominant cover type of even-aged stands with little evidence of replacement by shade-tolerant species.
Climax- The only tree species capable of growing in a particular environment; lodgepole pine is self-perpetuating.
Diseases and Parasites
Adult beetles attack lodgepole pine in July or August, introducing bluestain fungi (8). The beetles construct egg galleries in the phloem where larvae feed and together with the fungi, girdle and kill the tree. Larvae overwinter in the tree, complete development, and emerge as adult beetles in the spring.
Harvesting has been considered as a means of preventing mountain pine beetle epidemics (19). Silvicultural practices in an integrated program for controlling losses to mountain pine beetle have been suggested (9,18). No mortality occurred in heavily thinned stands in Oregon where vigor ratings were high (44).
The mountain pine beetle has played an historic role in the dynamics of lodgepole pine ecosystems. By periodically invading stands and creating large amounts of fuels, which are eventually consumed by fire, creating favorable conditions for regeneration (12,39), the beetle has increased the probability that lodgepole pine will reoccupy the site at the expense of other species.
Another aggressive bark beetle that attacks lodgepole pine is the pine engraver (Ips pini). Ips commonly develops in logging slash, especially slash that is shaded and does not dry quickly. Prompt slash disposal is an effective control measure. Ips also can build up in windthrows.
Other insects that can be damaging local pests are the lodgepole terminal weevil (Pissodes terminalis), which can be destructive to elongating terminal leaders; larvae of the Warren's collar weevil (Hylobius warreni), which girdles roots and the root collar; larvae of the weevil Magdalis gentilis, which mine branches; various sucking insects, such as the pine needle scale (Chionaspis pinifoliae), the black pineleaf scale (Nuculaspis californica), and the spruce spider mite (Oligonychus ununguis); and several defoliating insects, among which are the lodgepole sawfly (Neodiprion burkei), the lodgepole needle miner (Coleotechnites milleri), the sugar pine tortrix (Choristoneura lambertiana), the pine tube moth (Argyrotaenia pinatubana), and the pandora moth (Coloradia pandora) (7).
Dwarf mistletoe (particularly Arceuthobium americanum) is the most widespread and serious parasite affecting lodgepole pine (11,29). A. americanum seeds are forcibly ejected from the fruit for distances as great as 9 m (about 30 ft). The sticky seeds adhere to the foliage of potential host trees. The proportion of trees visibly infected can double each 5 years between the ages of 10 and 25, with nearly a third of the trees infected at age 25 (29).
Rate of spread in young stands is about 0.3 to 0.5 m (1.0 to 1.5 ft) per year, with the fastest rate in dense stands. In many areas, more than 50 percent of lodgepole pine forests are infected. Dwarf mistletoe infection results in reduced diameter and height growth, increased mortality, reduced wood quality, decreased seed production, and overall decreased vigor.
Both harvesting and fire can greatly lessen the rate of spread and rates of infection. Effective control can be accomplished by clearcutting and locating boundaries of the unit to minimize reinfection from surrounding stands. Fire can effectively limit spread of dwarf mistletoe by eliminating sources of infection and establishing vast acreages of dwarf mistletoe-free areas.
Lodgepole pine is subject to attack by many fungal pathogens (33). These fungi are responsible for reduced growth and considerable cull and mortality. They also contribute in no small measure to the large amounts of logging residues that commonly occur when lodgepole pine is harvested.
One of the most serious diseases in lodgepole pine is a stem canker caused by Atropellis piniphila. Cankered stems are usually useless for lumber or posts and poles. Stem cankers of rust fungi cause extensive mortality, growth loss, and cull in lodgepole pine. Of these comandra blister rust (Cronartium comandrae) isthe most serious. The western gall rust (Peridermium harknessii) is especially damaging; trunk cankers can cause cull in logs and can kill seedlings and saplings. Because this rust does not require an alternate host, it can directly reinfect pines. Other fungi attack lodgepole pine and may cause serious losses in wood production. Examples are needle casts (such as Elytroderma deformans and Lophodermella concolor); root rots (such as Armillaria mellea and Heterobasidion annosum); and wood decays (such as Phellinus pini and Peniophora pseudo-pini).
Seed and seedling diseases are not usually damaging, although locally several mold fungi are associated with seed losses in germination, and rotting and damping-off can affect young seedlings.
Because of its relatively thin bark, lodgepole pine is more susceptible to fire than Douglas-fir and many other associates. It is less susceptible than Engelmann spruce or subalpine fir. Mortality from beetle epidemics often creates large amounts of jackstrawed fuel, which ignites easily from lightning and other sources and hampers fire control efforts.
Chinook winds following extremely cold weather occasionally cause red belt injury, particularly in Canada and Montana. Defoliation of trees is common and mortality can occur over large areas. Heavy snow can break or bend trees, particularly in dense stands with narrow crowns and intense root competition. Thinning can contribute to snow breakage, particularly if previously dense stands are opened suddenly.
Animals can cause considerable damage in thinned stands in some areas. Porcupines were attracted to thinned and fertilized stands in Montana. Pocket gophers often cover small seedlings under their entrance mounds and "winter-casts." They also feed on or clip both roots and tops. Gopher populations often explode as vegetation increases in open areas.
Reaction to Competition
In the absence of fire, lodgepole pine is usually succeeded by its more tolerant associates, such as Engelmann spruce and subalpine fir. Succession proceeds at variable rates, however, and is particularly slow in some high elevation forests.
Pure stands of lodgepole pine persist for varying lengths of time. In northern Idaho and central Oregon, stands begin to break up at 80 to 100 years, while stands at higher elevations, such as in Montana, southern Idaho, Utah, and Wyoming, last for several hundred years. Pure stands in and around Yellowstone National Park contain 300- to 400-year-old trees, with several groups of younger even-aged trees. These stands no doubt originated as even-aged stands but have been breaking up for more than two centuries.
The ability of lodgepole pine to regenerate at the expense of other species is due not only to cone serotiny but also to seed viability, germinative energy, early rapid growth, and ability to survive a wide variety of microsite and soil situations (39).
Compared to its associates, lodgepole pine is intermediate in its needs for water, requiring more than Douglas-fir and ponderosa pine and less than Engelmann spruce and subalpine fir. On some sites, lodgepole pine appears to compete well for water, however, and grows where other species may be excluded because of lack of water (45,57); on others it appears to be tolerant of high water tables (14,43). It is also intermediate in its tolerance to extremes of temperature (27).
Lodgepole pine shows good response to thinning at an early age (17). Heavily stocked stands must be thinned before stagnation occurs. The best age for thinning varies with site and density. Poor sites and overstocked stands particularly must be thinned as early as age 10.
Diameter growth acceleration is usually greatest in heavy thinnings; cubic volume and basal area growth are usually greatest in light thinnings (27). Although mechanical thinning, as with bulldozer strips, is a convenient alternative, obtaining a proper response (36) is difficult.
At older ages, growth response is strongly correlated with crown size, vigor, and amount of release provided (27). Attempts at partial cutting of mature and over-mature stands have resulted in little gain or even negative net volume growth (1,28).
Lodgepole pine can be maintained best in a vigorous, productive forest by using a silvicultural method that regenerates even-aged stands (38). This often may be accomplished by clearcutting and by relying upon natural regeneration or planting. Planting provides an excellent opportunity for initial stocking control and/or genetic improvement.
Taproots and vertical sinkers are common, but where a hardpan or water table is encountered, the taproot may die, bend, or assume a horizontal position. Planting may affect root configuration. Taproots of seedlings planted with "J-roots" often grow horizontally for many years before sinkers develop.
Because of its shallow root system, lodgepole pine is susceptible to windfall, particularly after stands are opened by harvesting. Windfirmness varies with stand density, soil conditions, and topography. Shallow roots are common above hardpan or in shallow, rocky soils.
Life History and Behavior
Lodgepole pine cuttings are relatively easy to root. Adventitious roots have been developed artificially from 8-year-old lodgepole pine (by air-layering) after treatment with either indole-acetic or indole-butyric acid (17).
Callus tissue cultures and liquid cell suspensions have been produced from seedling hypocotyl tissue, excised embryos, and actively growing shoots.
Although lodgepole pine germinates well on most organic seedbeds, such materials tend to dry faster than mineral soil and seedlings often die in this seedbed. Lodgepole pine seeds have little need for stratification and germination depends largely upon temperature (20). At optimum temperatures and moisture, almost 100 percent of the seeds germinate rapidly.
Both shading and competition inhibit germination and survival. Newly germinated seedlings are relatively insensitive to temperature extremes. Because residual overstory following partial cutting usually does not provide the most favorable conditions for regeneration, clearcutting is generally recommended. On some areas, however, lodgepole pine has established itself in the shade of lightly cut or uneven-aged stands and may persist for many years in the understory. Some of these trees eventually may establish a crown sufficient to permit reasonable growth.
Drought is a common cause of mortality among first-year seedlings; losses vary with soil type and seedbed condition. Greatest losses occur on soils with low water-holding capacity, and duff and litter. Well decomposed organic material, incorporated in the soil, enhances seedling survival, however. Disturbed mineral soil seedbeds generally produce the best germination and survival (34,40,41). Shading has been demonstrated to help under drought conditions in Wyoming (10).
Drought losses usually decline considerably after the first growing season. First-year seedlings are particularly vulnerable because of a relatively shallow root system (34,47).
Young, succulent seedlings may die because of high soil surface temperatures (13). By 2 to 4 weeks of age, seedlings are able to withstand soil surface temperatures higher than 60° C (140° F), which commonly occur at high elevation sites. Freezing temperatures may kill seedlings either directly or by frost heaving. In much of the range of lodgepole pine, however, frosts occur regularly throughout the growing season and seedlings from different sources vary in frost resistance (16). The amount of frost heaving varies considerably by soil type, location, and year of occurrence but can cause significant losses.
Lodgepole pine seedlings are poor competitors and competition from grass is often most detrimental. The Douglas-fir/pinegrass habitat type is one of the most difficult sites for lodgepole pine regeneration, particularly if the regeneration effort is delayed until a firm sod cover is established.
Grazing animals, particularly cattle, can cause seedling mortality by trampling. Sheep actually seek the succulent new "candles" in the spring and nibble needles and small branches if other feed is not abundant.
A common problem of regenerating lodgepole pine stands is overstocking, which results in stagnation at early ages. Many sites are stocked with tens of thousands and even hundreds of thousands of trees per hectare.
If trees are well distributed, stocking should not exceed 1,240 to 1,980 stems per hectare (500 to 800/acre) between 5 years and 20 years of age (17). Proper distribution and full utilization of the site, however, may require establishment of 2,470/ha (1,000/acre) and thinning to obtain proper spacing. There is also potential for significant genetic gains from selection of elite trees when thinning.
An average height of 3.6 m (12 ft) and d.b.h. of 5 cm (2 in) on fully stocked 20-year-old stands was found on above average sites in Montana (27). Average heights of 2.0 m (6.7 ft), 4.2 m (13.8 ft), and 7.6 m (24.9 ft) were found on low, medium, and high sites in 20-year-old stands in the Foothills Section of Alberta (for density class 1,240 stems per hectare or 500/acre at 70 years of age) (32).
Lodgepole pine height growth begins earlier than any of its associates except other pines and larch (53).
Seed Production and Dissemination
Lodgepole pine is a prolific seed producer. Good crops can be expected at 1- to 3-year intervals, with light crops intervening. The cones withstand below freezing temperatures and are not generally affected by cone- and seed-feeding insects. Only squirrels and coreid bugs are significant seed predators. Seed production should not be taken for granted, however. Complete seed crop failures have occurred at 2800 m (9,200 ft) in northwest Wyoming for 2 to 4 years in a row (42).
Cone production of individual dominant and codominant trees can vary from a few hundred to a few thousand per tree (37). Cones are persistent, and serotinous (closed) cones accumulate for decades. Annual production may run from 173,000 to 790,000 seeds per hectare (70,000 to 320,000/acre) with half to one-third available for annual seedfall (27). An annual seedfall of 99,000 to 222,000 seeds per hectare (40,000 to 90,000/acre) was found in central Montana (58). These figures might be considered typical for interior lodgepole pine where some portion of the trees are of the serotinous type. In Oregon, where the nonserotinous cone habit is prevalent, seedfall ranged from about 35,000 to over 1.2 million/ha (14,000 to 500,000/acre) (21). Most years seedfall was on the order of hundreds of thousands per hectare. Where stored seeds are in the millions per hectare (in closed cones), the number of seeds stored is probably 10 times that of seeds produced annually (37).
Although the number of fully developed seeds per cone varies from as few as 1 to 2 to as many as 50, a normal average for large cone lots in the Rocky Mountains is from 10 to 24 seeds per cone (42). Sierra Nevada populations range from 5 to 37 seeds per cone (20).
The serotinous cone habit varies over wide geographic areas as well as locally (37). Serotinous cones are not common in eastern Oregon, rare in coastal populations, and absent in the Sierra Nevada and southern California and Baja California populations (20). Although common in the Rocky Mountains, this cone habit varies considerably (37). Many stands in the Rockies have less than 50 percent serotinous-cone trees.
Lodgepole pine has long been regarded as a fire-maintained subclimax type. Its ability to regenerate in extremely dense stands to the exclusion of other species can be attributed to the closed cone habit. Millions of seeds per hectare are held in reserve for many years and are readily available to germinate on the seedbed prepared by fire. Recent evidence seems to indicate that fire selects strongly for the closed cone habit (49).
Serotinous cones do not open at maturity because of a resinous bond between the cone scales. The bonds break with temperatures between 45° and 60° C (113° to 140° F) (48), and cone scales are then free to open hygroscopically. Large quantities of seeds are thus available for regenerating a stand following fire. Closed cones at or near the soil surface (less than 30 cm or about 12 in) are also subjected to temperatures from insolation sufficient to open them and may provide seed in harvested areas. Some seeds may be damaged by fire, however, particularly in fires burning in logging slash.
Seeds stored in serotinous cones on the tree remain viable for years. Apparently, prolonged viability can be maintained so long as cones or seeds are not in contact with the ground. Once cones are on the ground, cones open. Damping-off fungi may infect the seed, rodents may feed on the seeds, or germination may occur; for the most part, seeds are not stored in the soil.
Lodgepole pine has relatively small seeds for pine. Seed weights vary considerably, ranging from 2.3 mg (0.04 grains) per seed in the Interior of Canada to 11.4 mg (0.18 grains) per seed in the Sierra Nevada (20). Lodgepole pine seeds average about 298,000 cleaned seeds per kilogram (135,000/lb) for varieties contorta, 258,000/kg (117,000/lb) for murrayana, and 207,000/kg (94,000/lb) for latifolia (54). Density of seedfall 20 m (66 ft) from the timber edge is only 10 to 30 percent of that at the timber edge for stands in the Rocky Mountains (fig. 1) (42). Dispersal of sufficient seed to adequately restock an area often is only about 60 m (200 ft) (23,38). Prevailing winds, thermal effects, or scudding on the snow may disperse seeds far beyond these distances, however.
Figure 1- Sound seed per hectare as a function of distance
from the nearest timber edge.
The annual seedfall from nonserotinous cones helps in restocking relatively minor disturbances in the stand, in maintaining the presence of lodgepole pine in mixed stands, and in expanding conifers into other vegetative types. Seldom do we find stands without some trees of the open-coned type. The efficacy of this seed source can be seen in the dense stands of lodgepole pine along road cuts, powerline rights-of-way, and ditches or where disturbance occurs near lodgepole pine stands.
Studies of seedfall have shown variation in the number of seeds released soon after cone maturation, but most seeds (80 to 90 percent) are released before the following growing season (27).
Where large amounts of seed are stored in serotinous cones, a most effective means of seed dispersal in clearcuts is from cones attached to the slash and those knocked from the slash and scattered over the forest floor during slash disposal. Many cones on or near the ground are opened by normal summer soil surface temperatures (35). In Montana 83 percent of the cones on the ground opened the first year on south slopes compared to 40 percent on north slopes. Maximum seed release from serotinous cones near the ground takes place during the first year of exposure. In fact, cones may open after the first few minutes of exposure to temperatures high enough to break the resinous bonds.
In slash, serotinous cones that are well above the ground behave like those on a tree- they remain closed, and stored seeds remain viable for years.
Seeds in unopened cones and those released from the slash may also be lost to rodents, fungi, and other destructive agents. Seeds from closed cones are usually available only for the first growing season following harvest, but stocking from open-cone seed sources can continue to increase for several years.
Slash disposal on areas where regeneration is planned from serotinous cones must be carefully planned and executed. Seed supply will be largely destroyed if slash to be burned is piled before cones have had a chance to open (38). Piling slash should be delayed until sufficient cones have opened to assure adequate stocking. Piling then scatters seeds and opened cones and helps prepare the seedbed. Piling slash after germination can also decrease stocking because young seedlings are trampled or buried.
Broadcast burning may hasten release of seeds from cones not in a position to open from high soil-surface temperatures. Some seeds will be destroyed, however; the amount will vary with fire intensity.
Flowering and Fruiting
Pollen generally matures in mid-May to mid-July (table 1) (20,52). The time at which pollen matures appears to be related to elevation and climate.
Table 1- Time of pollen shedding in natural stands of lodgepole pine (20,52, modified)
Elevation¹ Years observed Date of peak shedding m ft Vancouver, BC - - 2 Middle to late May Northwestern Washington 150 500 10 May 12 Mendocino White Plains, California - - 1 June 9 Northern Cascades 1200 4,000 - Mid-June Northern Idaho; western Montana - - 10 June 13 Central and eastern Washington and Oregon 790 to 1300 2,600 to 4,250 - June 13 Southeastern Alberta (subalpine forest) - - 10 June 22 Sierra Nevada, California 1820 6,000 3 June 22 Central Montana; Yellowstone region - - 10 June 25 Northern Utah 2190 7,200 2 July 12 Southern Idaho 2070 6,800 1 July 7 Northern Idaho; western Montana 670 to 1265 2,200 to 4,150 10 June 6 Eastside Montana; Yellowstone National Park 975 to 2060 3,200 to 6,750 10 June 17 ¹Dash indicates data are not available. Seed cones usually mature in August, September, or October, more than a year after pollination. Inland forms and high elevation stands apparently mature earlier than coastal forms or low elevation stands. Cones open in early September in the Northern Rocky Mountains. Cone maturity is indicated by a change in color from purple-green to light brown (54).
Growth and Yield
Maximum yield in the Rocky Mountains was 280 m³/ha (20,000 fbm/acre) at a density of 1,980 trees per hectare (800/acre), but only 21 m³/ha (1,500 fbm/acre) at a density of 4,450/ha (1,800/acre), assuming 5 fbm/ft³; original figures were in board feet (27).
In extreme cases 70-year-old stands with 247,000 trees/ha (100,000 trees/acre) averaged only 1.2 m (4 ft) in height and less than 2.5 cm (1 in) in diameter at ground level.
Yields of 168 to 224 m³/ha (about 12,000 to 16,000 fbm/acre) can be found in old-growth Rocky Mountain lodgepole pine. Yields of more than 336 m³/ha (about 24,000 fbm/acre) are the result of a fortuitous combination of favorable initial stocking, good site quality, and absence of mountain pine beetle and dwarf mistletoe.
Relationships among age, stocking levels, and development in natural stands were summarized for medium sites in Montana and Idaho (site index 22.9 m or 75 ft at 100 years) (table 2). Under light to moderate stocking, live crowns are 25 to 60 percent of total height.
Table 2- Relationships among stand age and stocking level, and tree development and typical yield in natural stands of lodgepole pine, summarized for medium sites in Montana and Idaho (site index 22.9 m or 75 ft at base age of 100 years)¹
Stocking Average height of dominants Average stand diameter
Total cubic volume
Merchantable volume yr trees/ha trees/acre m ft cm in m³/ha ft³/acre m³/ha ft³/acre fbm/acre 20 1,240 500 5.5 18 8.6 3.4 16.1 230 - 19,770 8,000 3.0 10 4.1 1.6 28.0 400 - 50 1,180 479 12.5 41 16.5 6.5 144.9 2,070 130.2 1,860 5,100 15,200 6,150 9.1 30 6.9 2.7 165.9 2,370 - - - 80 1,030 418 18.0 59 20.6 8.1 285.6 4,080 266.0 3,800 12,100 7,500 3,034 14.6 48 9.1 3.6 280.0 4,000 - - - 110 850 344 22.3 73 23.6 9.3 385.7 5,510 363.3 5,190 18,200 4,600 1,861 18.9 62 11.4 4.5 357.0 5,100 273.0 3,900 8,400 140 680 275 25.3 83 26.7 10.5 448.7 6,140 426.3 6,090 23,200 3,070 1,243 22.3 73 14.0 5.5 416.5 5,950 301.0 4,300 10,300 ¹Compiled from unpublished yield tables furnished by D.M. Cole, USDA Forest Service, Intermountain Research Station, Bozeman, MT. Cubic volumes are from trees 11.4 cm (4.5 in) in d.b.h. to a 7.6 cm (3 in) diameter top. Board foot volumes are from trees larger than 16.5 cm (6.5 in) in d.b.h. to a 15.2 cm (6 in) diameter top. Mature sizes vary greatly between stands. In the Rocky Mountains, most trees at 140 years of age were 18 to 33 cm (about 7 to 13 in) in d.b.h. and 18 to 25 m (about 60 to 80 ft) in height (27).
Trees in the Blue Mountains of Oregon average 30 cm (about 12 in) in d.b.h. and 23 m (about 75 ft) tall at 100 years of age. Sierra Nevada trees the same age are larger, averaging 42 cm (about 16 to 17 in) in d.b.h. and 28 to 30 m (about 90 to 100 ft) tall. Coastal trees are smaller but vary greatly. Mature trees range from 15 to 50 cm (about 6 to 20 in) in d.b.h. and only 6 to 12 m (20 to 40 ft) tall. Dwarf lodgepole pines are only about a meter (2 to 5 ft) tall and are found along the coast in Mendocino County, CA. This small size is thought to be caused by a highly acid hardpan.
Growth of lodgepole pine is often so stagnant that stand culture is not practical. Early management and control of stocking greatly affects growth and yield of lodgepole pine stands (17). Average annual growth in old-growth unmanaged stands in the central Rocky Mountains only was 0.4 to 0.6 m³/ha (about 25 to 40 fbm/acre) because of large numbers of small trees and a high incidence of dwarf mistletoe (4). (Calculations assume 5 fbm/ft³; original figures were in board feet). Annual net growth may be increased to 2.1 to 5.6 m³/ha (about 150 to 400 fbm/acre) by controlling stand density and reducing dwarf mistletoe infection (5,25).
Control of stand density offers the greatest opportunity for increasing productivity of any readily available management practice.
Culmination of total cubic volume occurs as early as 40 years in severely stagnated stands, and between 50 and 80 years for overstocked, but not greatly stagnated, stands. Merchantable volume culmination in stands of the latter type occurs between 110 and 140 years, depending on merchantability standards.
Thinning of young overstocked and stagnating stands can restore growth potential and redirect it into merchantable-size products. With more complete utilization (lower merchantable d.b.h. and top diameters), most of the yield increase possible from thinning is attained with the first entry, a stocking control thinning (17).
Molecular Biology and Genetics
Barcode data: Pinus contorta
Statistics of barcoding coverage: Pinus contorta
Public Records: 8
Specimens with Barcodes: 17
Species With Barcodes: 1
IUCN Red List Assessment
Red List Category
Red List Criteria
National NatureServe Conservation Status
Rounded National Status Rank: N5 - Secure
Rounded National Status Rank: N5 - Secure
NatureServe Conservation Status
Rounded Global Status Rank: G5 - Secure
Reasons: Extensive range from Alaska to northern New Mexico and Baja California, attaining best populations in the Rocky Montains (Record and Hess 1943). Thrives in mostly well drained soils but may be found in peat bogs, muskegs or on dry sandy sites. Successfully cultivated in New Zealand and the United Kingdom (Tree Talk 1994).
Comments: The timber is well known (Tree Talk 1994).
Relevance to Humans and Ecosystems
Uses: FIBER, Building materials/timber
Comments: Uses of the timber are local for mining timbers and railway crossties, sometimes under the name of Tamarack (Record and Hess 1943). Other common applications are corral rails, veneer, hardboards, mine timbers, orchard props, paneling, particleboard, plywood, poles, posts, pulpwood, rough construction, rustic furniture, shingles and siding (Tree Talk 1994).
Lodgepole pine is used for framing, paneling, posts, corral poles, utility poles, railroad ties, and pulpwood. As new developments such as structural particleboard become practical, the rapid juvenile growth of the species will be an advantage where gross cubic volume becomes important. Even now, with properly designed machinery, it is economically harvested, and this harvesting, properly done, can enhance watershed, forage, wildlife habitat, and scenic and recreational values.
Pinus contorta, with the common names lodgepole pine and shore pine, and also known as twisted pine, and contorta pine, is a common tree in western North America. It is common near the ocean shore and in dry montane forests to the subalpine, but is rare in lowland rain forests. Like all pines (member species of the genus Pinus), it is an evergreen conifer.
- Pinus contorta subsp. bolanderi — Bolander's beach pine, Bolander pine; endemic to NW California Coast (e.g. Mendocino County); Near Threatened by fires and development
- Pinus contorta subsp. contorta — Shore pine; Pacific Coast, southern Alaska to California.
- Pinus contorta subsp. contorta var. contorta — Shore pine; Pacific Coast, Northwest California through Alaska.
- Pinus contorta subsp. murrayana — Tamarack pine, or Sierra lodgepole pine; Cascade Range from Washington into Northern California; the Sierra Nevada, the Transverse Ranges of Southern California (including the San Bernardino Mountains, the Peninsular Ranges into northern Baja California, and the Spring Mountains of southern Nevada).
- Pinus contorta subsp. latifolia — Lodgepole pine; Rocky Mountains, Colorado to Yukon and Saskatchewan; Aspen parkland and boreal forests.
Depending on subspecies, Pinus contorta grows as an evergreen shrub or tree. The shrub form is krummholz and is approximately 1 to 3 m (3.3 to 9.8 ft) high. The thin and narrow-crowned tree is 40 to 50 m (130 to 160 ft) high and can achieve up to 2 m (6.6 ft) in diameter at chest height. The murrayana subspecies is the tallest. The crown is rounded and the top of the tree is flattened. In dense forests, the tree has a slim, conical crown. The formation of twin trees is common in some populations in British Columbia. The elastic branches stand upright or overhang and are difficult to break. The branches are covered with short shoots that are easy to remove.
The species name is contorta because of the twisted, bent pines found at coastal areas and the tree's twisted needles. Pinus contorta is occasionally known under several English names: black pine, scrub pine, and coast pine. P. contorta subsp. latifolia will hybridise with the closely related Jack pine (Pinus banksiana).
Needles and buds
The egg-shaped growth buds are reddish-brown and between 20 and 30 mm (0.79 and 1.18 in) long. They are short pointed, slightly rotated, and very resinous. Spring growth starts in beginning of April and the annual growth is completed by early July. The dark and mostly shiny needles are pointed and 4 to 8 cm (1.6 to 3.1 in) long and 0.9 to 2 mm (0.035 to 0.079 in) wide. The needle edge is weak to clearly serrated. The needles are in pairs on short shoots and rotated about the shoots' longitudinal axes. In Alberta above 2,000 m (6,600 ft), 1 to 5 needles occur per short shoot. A population with a high proportion of three-needled short shoots occurs in the Yukon. Needles live an average of four to six years, with a maximum of 13 years.
The 3–7 cm cones often need exposure to high temperatures (such as from forest fires) in order to open and release their seeds, though in subsp. murrayana they open as soon as they are mature. The cones have prickles on the scales.
Pinus contorta is a fire-dependent species, requiring wildfires to maintain healthy populations of diverse ages. The bark of the lodgepole pine is fairly thin, minimizing the tree's defense to fire; however, the heat of these closed-cone pine forest rejuvenating fires open the cones to release the seeds. This allows the species to regenerate and maintain its place in the forest habitat.
Excessive wildfire prevention disrupts the fire ecology. The stands are usually so densely populated that the trees self-thin, or out-compete each other, leaving dead trees standing. These become a dry ladder fuel that can accelerate the fire to the crown of living trees. When the fire reaches the crowns of the trees, it can jump from tree to tree and becomes relatively unstoppable.
The natural fire regime for this species is primarily driven by climate. The fires occur most often after years of drought. Pinus contorta occurs from the upper montane to the subalpine region. These types of forests experience a lot of moisture in the form of snow in the winter due to their altitude. The density of the tree stand also prohibits the establishment of an understory. With all of that being said, the likelihood of a surface fire occurring is rare. Thus, infrequent but severe fires dominate this species.
An example of the climate that plays a huge role in the fire regime of Pinus contorta is quite complex. There are three different oscillations that play a major role in droughts. These are the Pacific Decadal Oscillation (PDO), Atlantic Multi-decadal Oscillation (AMO) and El Nino (ENSO). A combination of these oscillations being in effect (+) or not in effect (-) have a global effect on the water available to these forests. So when the AMO +, ENSO – and PDO -, there is going to be a drought and likely a severe subalpine fire.
Suillus tomentosus, a fungus, produces specialized structures called tuberculate ectomycorrhizae with the roots of lodgepole pine (Pinus contorta var. latifolia). These structures have been shown to be the location of concentrations of nitrogen-fixing bacteria which contribute a significant amount of nitrogen to tree growth and allow the pines to colonize nutrient-poor sites.
Native American tipis
Lodgepole pine is named for its common use as structural poles for the Native American tipi shelter. A typical tipi is constructed using 15 to 18 pines. The long, straight, and lightweight characteristics of the species made it ideal for horse transport in nomadic Plains buffalo hunting cultures. Tribes made long journeys across the Great Plains to secure lodgepole pines that only grew in mountainous areas. In Minnesota, other species such as red pine would be used in tipis, though they were generally thicker, heavier, and more cumbersome to transport than Pinus contorta.
Pinus contorta is still used by many today for erecting tipis on American Indian reservations, at powwows, and at private homes. The trees may be harvested for tipi poles in U.S. National Forests, provided the harvester secured a permit to cut living trees for ceremonial or traditional purposes. The Bighorn Mountains, the Black Hills, and the Medicine Bow Mountains are popular tipi pole harvesting areas for Native Americans living on Plains Indians reservations in North and South Dakota, and immigrant tipi enthusiasts.
Pinus contorta is cultivated as an ornamental tree by the horticulture industry. Plant nurseries grow Pinus contorta subsp. contorta and Pinus contorta subsp. murrayana for use in traditional and wildlife gardens, and as smaller selections of the native plant for natural landscaping. The Shore pine's (ssp. contorta) smaller varieties and cultivars are also used in container gardening, including as large bonsai specimens.
Cultivars of this species include:
- Chief Joseph, a dwarf variety of Pinus contorta var. latifolia grown for its yellow winter needles.
- Spaan's Dwarf, a dwarf variety of Pinus contorta var. contorta that grows wider than it grows tall.
Lodgepole pine is the Provincial tree of Alberta, Canada.
Pinus contorta is a serious invasive species of wilding conifer in New Zealand, along with several other western North American pine species. It is listed on the National Pest Plant Accord and is prohibited from sale, commercial propagation, and distribution.
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