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General DescriptionTrees to 20 m tall; trunk to 30 cm d.b.h.; bark dark gray or gray, fissured. Branchlets purplish brown, pubescent, sometimes stipitate glandular, glabrescent, with scattered, minute lenticels. Petiole 1-3.5 cm, densely pilose when young, glabrescent; leaf blade ovate-oblong or obovate-oblong, 5-15 cm long, 3-9 cm wide, papery, abaxially bearded in axils of lateral veins, pilose and sparsely glandular, adaxially sparsely pilose especially along impressed midvein when young, base very obliquely rounded or subcordate, margin sharply and doubly mucronate serrate, apex long acuminate to caudate-acuminate; lateral veins 10-14 on each side of midvein. Male inflorescence ca. 2 cm; bracts broadly ovate, tomentulose, apex acute to acuminate; anthers purple. Female flowers 4-6 in a cluster; bracts campanulate, densely pubescent, sometimes stipitate glandular, with laciniate segments forming branching, pungent spines. Nut ovoid-globose, subcompressed, 1-1.5 cm in diameter, apex pubescent.