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BiologyOccurs in lagoons and lakes, large- to medium- size rivers, on sand or mud bottom; usually on open sand or sand-shell bottom. Lives up to 3 years. Spawns for the first time at 1 year, rarely at 2. Spawns in April-September. Individual females may repeat spawning during a season. Adhesive eggs are deposited on stones, shells and aquatic plants and are guarded by males until hatching. Feeds on a wide variety of invertebrates (mainly crustaceans and insect larvae) and small fish (Ref.59043). Is the most important commercial species of goby in the Caspian Sea (Ref. 2058).