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DescriptionAdult Lt 635–657 µm; L to PhJIn 155–163 µm at U26–U23 (Fig. 17). Body elongate, slender; mouth a narrow outwardly rolled protrusion, head with a band of circumcephalic cilia at U03 and prominent pestle organs at U04, but lacking lateral lobes; neck constriction lacking, body sides parallel over most of their length, thinning gradually to the caudal base; caudum is slightly cleft, incised from its tips to U97; medial cone is usually absent. Widths at narrowed mouth /pestle organs /PhJIn /mid-trunk /furcal base, tips, and their locations along the body length are: 18 /31 /38 /44 /18, 30 µm at U02 /U04 /U23 /U58 /U97, U100. Epidermal glands ca. 28 per side, small (2–5 µm diameter), are distributed along the lateral body margins, but appear inconspicuous. Adhesive tubes: TbA 8 per side (L 5–9 µm), the shortest one inserting on the medial most edge, occur on fleshy hands that insert at U11; TbL absent; TbD 7 per side (L 11–14 µm) from U27 to U84 and TbV 14 per side (L also 11–14 µm) from U29 to U88, all of similar size and spacing in the intestinal region; ‘dohrni’ [Seitenfüsschen] tubes 2 per side, posteriolaterally directed (L longer tube 20 µm, shorter 15 µm), inserting ventrolaterally immediately behind the fleshy hands at U12; TbP 8 per side, the outermost being the longest and thickest (L 14 µm), the others being shorter (L 3–9 µm), none occurring on the lateral or leading edges of the lobes. Ciliation: Head protrusion has sensory hairs (L 11–30 µm) laterally and a circumcephalic band of cilia (L 14 µm) at U03; other sensory hairs (L 20–30 µm) occur on the trunk in lateral, dorsolateral and dorsal columns, with 14/14/18 per side. Ventral locomotor cilia (L=8 µm) flow from the circumcephalic band rearward in two longitudinal bands that trace the lateral body margins, joining again behind the level of the anus. Digestive tract: Mouth terminal, narrow (8 µm diameter); buccal cavity small, deep, vaseshaped; walls of medium cuticularization; basal pharyngeal pores are large and conspicuous, but lack pharyngeal knobs; intestine broadest in front, narrowing gradually to the rear, with a bulge at the level of the ventral anus, U93. Reproductive tract: Hermaphroditic, protandrous to simultaneous; paired testes extend rearward from just behind the PhJIn, their vasa deferentia recurving from the rear to the fore and exiting behind together at U31; small developing ova (61 × 13 and 41 × 8 µm) occur bilaterally in the mid-gut region; neither frontal nor caudal organs were seen.