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BiologyLacustrine, pelagic, in upper water layers during summer and descending to deeper layers in winter. It spawns at 1-2 years, near shores above gravel or coarse sand, with most individuals spawning more than one season. Spawning starts when temperature reach about 19-20°C, in July to August (Ref. 59043). Adults form schools in the epilimnion during summer and tend to congregate in shallow waters near shores. They feed on plankton (cladocerans and copepods) and small fishes (Ref. 10545, 59043). It is threatened by water abstraction and pollution (Ref. 26100).