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BiologyInhabit inshore waters (Ref. 3167). Depth range to 170 m (Ref. 04426), and from 333-343 m in the eastern Ionian Sea (Ref. 56504). Feed on all kinds of benthic animals (Ref. 3167). Oviparous. Distinct pairing with embrace. Young may tend to follow large objects, such as their mother (Ref. 205). Eggs are oblong capsules with stiff pointed horns at the corners deposited in sandy or muddy flats (Ref. 205). Egg capsules are 4.5 cm long and 3.0 cm wide (Ref. 41304).