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BiologyAdults are found in the middle reaches of rivers during the summer; migrates downstream in autumn (Ref. 559). River bed consists of cobbles, boulders and gravel in riffles and sand and gravel in pools (Ref. 56353). Feed on fishes (Ref. 559). Juveniles migrate from the spawning ground in the sea to the middle reaches of rivers to grow (Ref. 56353).