The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • Beal, R. S., Jr. 2003. Annotated Checklist of Nearctic Dermestidae with Revised Key to the Genera. The Coleopterists' Bulletin, vol. 57, no. 4
  • Beal, R. S., Jr. 2003. Annotated Checklist of Nearctic Dermestidae with Revised Key to the Genera. The Coleopterists' Bulletin, vol. 57, no. 4. 391-404
  • CITES. Convención sobre el comercio Internacional de Especies Amenazadas de Fauna y Flora Silvestres. 2011. Apéndices I, II y III. Fecha de actualización: 14 de octubre 2010.
  • Ciro, D. W., Anteparra, M. E. & Hermann, A. 2008. Dermestidae (Coleoptera) en el Perú: revisión y nuevos registros. Rev. Per. Biol. 15(1): 15-20.
  • DAISIE (Delivering Alien Invasive Species Inventories for Europe). 2011. Delivering Alien Invasive Species Inventories for Europe.
  • Ellis, A. M. & Hodges, A. C. 2007. Handout developed to accompany the Khapra Beetle Trogoderma granarium Everts (Coleoptera: Dermestidae) NPDN First Detector Training Module. University of Florida, SPDN
  • French, S. & Venette, C. R. 2005. Mini Risk Assessment  Khapra Beetle, Trogoderma granarium (Everts)  [Coleoptera: Dermestidae]. CAPS PRA
  • GISD (Global Invasive Species Database). 2007. Trogoderma granarium. UICN. Consultada 28 de noviembre 2011 en
  • Hinton, H. 1945. A monograph of the beetles associated with stored products. British Museum Vol I. Jarrold and Sons Ltd. Londo: 443
  • ISC (Invasive Species Compendium). 2011. Trogoderma granarium. Consultado el 24 de noviembre de 2011 en CABI
  • ISSG. 2008. Global Invasive Species Database. IUCN. March
  • IUCN. 2011. IUCN Red list of threatened species, Version 20011.2.
  • Karnowski, W. 2002. Trogoderma granarium, diagnostic protocols for regulated pests. Bulletin OEPP/EPPO Bulletin. 32: 299-301
  • Lepesme, P. 1994. Les coleopteres des denrées alimentairies et des produits industriets entreposés. Encyclopedie Entomologique. Serie A, XXII Ruede Tournon, Paris: 335
  • Lowe, S., Browne, M., Boudjelas, S., De Poorter, M. 2000. 100 of the World´s Worst Invasive Alien Species, a selection from the Global Invasive Species Database. The Invasive Species Specialist Group of the Species Survival Commission (SSC) of the World Conservation Union (UICN). 12 pp. Consultado en
  • NIMF 27. 2006. Trogoderma granarium Everts (2012). Anexo 3. Roma, CIPF, FAO.
  • Orrell T. (custodian) (2015). ITIS Regional: The Integrated Taxonomic Information System (version Dec 2014). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
  • Pasek, J. E. 1998. Khapra beetle (Trogoderma granarium Everts): pest-initiated pest risk  assessment. USDA APHIS. Raleigh, NC 32. pp.
  • Rebolledo, R. & Arroyo, M. 1995. Studies of behaviour of Trogoderma granarium Everts (Coleoptera: Dermestidae) in diapause owed to the absence of food. Bol. San. Veg. Plagas. 21(3): 319-327
  • SEMARNAT. 2010. NORMA Oficial Mexicana NOM-059-SEMARNAT-2010, Protección ambiental-Especies nativas de México de flora y fauna silvestres-Categorías de riesgo y especificaciones para su inclusión, exclusión o cambio- Lista de especies en riesgo. Diario Oficial de la Federación. 30 de diciembre de 2010. D.F., México. 85 p
  • SENASA. 2000. Monitoreo preventivo del Gorgojo khapra (Trogoderma granarium Everts.). Manual del sistema de trampeo. Ministerio de Agricultura. Lima, Perú
  • United Nations Environment Programme. 2000. The Montreal protocol on substances  that deplete the ozone layer as either adjusted and/or amended in London 1990,  Copenhagen 1992, Vienna 1995, Montreal 1997, Beijing 1999. Secretariat for the  Vienna Convention for the Protection of the Ozone Layer


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