The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • Barton, B., J. Roth. 2008. Implications of intraguild predation for sea turtle nest protection. Biological Conservation, 141:8: 2139-2145.
  • Branco, J., J. Hillesheim, H. Fracasso, M. Christoffersen, C. Evangelista. 2010. Bioecology of the ghost crab Ocypode quadrata (Fabricius, 1787) (Crustacea: Brachyura) compared with other intertidal crabs in the Southwestern Atlantic. Journal of Shellfish Research, 29 (2): 503-512. Accessed June 06, 2011 at
  • Burggren, W. 1988. Biology of the Land Crabs. New York, NY: Cambridge University Press.
  • Diaz, H., J. Costlow. 1972. Larval development of Ocypode quadrata (Brachyura: Crustacea) under laboratory conditions. Marine Biology, 15: 120-131. Accessed June 06, 2011 at
  • Felder, D. L., F. Álvarez, J. W. Goy, and R. Lemaitre. 2009. Decapoda (Crustacea) of the Gulf of Mexico, with Comments on the Amphionidacea,
  • Fisher, J., M. Tevesz. 1979. Within-habitat spatial patterns of Ocypode quadrata (Fabricius) (Decapoda Brachyura). Crustaceana, Supplement No. 5: 31-36.
  • Gosner, K. L. (1978). A Field Guide to the Atlantic Seashore. Boston: Houghton Mifflin.
  • Haley, S. 1969. Relative growth and sexual maturity of the Texas ghost crab, Ocypode quadrata (Fabr.) (Brachyura, Ocypodidae). Crustaceana, 17 (3): 285-297.
  • Haley, S. 1972. Reproductive cycling in the ghost crab, Ocypode quadrata (Fabr.) (Brachyura, Ocypodidae). Crustaceana, 23 (1): 1-11.
  • Hobbs, C., C. Landry, J. Perry. 2008. Assessing anthropogenic and natural impacts on ghost crabs (Ocypode quadrata) at Cape Hatteras National Seashore, North Carolina. Journal of Coastal Research, 24 (6): 1450-1458.
  • Knott, D. 2010. "Atlantic ghost crab: Ocypode quadrata" (On-line). Accessed June 06, 2011 at
  • McDermott, J. 2009. Notes on the unusual megalopae of the ghost crab Ocypode quadrata and related species (Decapoda: Brachyura: Ocypodidae). Northeastern Naturalist, 16 (4): 637-646.
  • Mitchell, P. 2007. "Ghost Crab: hungry nocturnal ghosties" (On-line). Mitchells Publications. Accessed June 06, 2011 at
  • Ng P.K.L. & Davie P. (2015). WoRMS Brachyura: World List of marine Brachyura (version 2015-08-01). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
  • Nizinski, Martha S. 2003. Annotated checklist of decapod crustaceans of Atlantic coastal and continental shelf waters of the United States. Proceedings of the Biological Society of Washington, vol. 116, no. 1. 96-157
  • Portell, R., R. Turner, J. Beerensson. 2003. Occurance of the Atlantic ghost crab Ocypode quadrata from the Upper Pleistocene to Holocene Anastasia formation of Florida. Journal of Crustacean Biology, 23 (3): 712-722.
  • Rothschild, S. 2004. Beachcomber’s Guide to Gulf Coast Marine Life: Third Edition: Texas, Louisiana, Mississippi, Alabama and Florida. Lanham, MD: Taylore Trade Publishing.
  • Schlacher, T., S. Lucrazi. 2009. Monitoring beach impacts: a case for ghost crabs as ecological indicators?. 2nd Queensland Coastal Conference, Gold Coast: 1-15.
  • Shields, J. 1998. "The ghost crab, Ocypode quadrata" (On-line). Virginia Institute of Marine Science. Accessed June 06, 2011 at
  • Williams, Austin B., Lawrence G. Abele, D. L. Felder, H. H. Hobbs, Jr., R. B. Manning, et al. 1989. Common and Scientific Names of Aquatic Invertebrates from the United States and Canada: Decapod Crustaceans. American Fisheries Society Special Publication 17. 77
  • Wolcott, T. 1978. Ecological role of ghost crabs, Ocypode quadrata (Fabricius) on an ocean beach: scavengers or predators?. Journal of Experimental Marine Biology and Ecology, 31 (1): 67-82.
  • da Rosa, L., C. Borzone. 2008. Spatial distribution of the Ocypode quadrata (Crustacea: Ocypodidae) along estuarine environments in the Paranagua Bay Complex, southern Brazil. Revista Brasileira de Zoologia, 25 (3): 383-388. Accessed June 06, 2011 at


EOL content is automatically assembled from many different content providers. As a result, from time to time you may find pages on EOL that are confusing.

To request an improvement, please leave a comment on the page. Thank you!