The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • Amphibian Species of the World V2.20 Database. The American Museum of Natural History. Department of Herpetology.">">
  • Bogart, J. P. (1991). ''The influence of life history on karyotypic evolution in frogs.'' Amphibian Cytogenetics and Evolution. D.M. Green and S.K. Sessions, eds., Academic Press, San Diego, 233-258.
  • Costa Rica, Asamblea Legislativa. Decreto No. 26435-MINAE. La Gaceta, Diario Oficial (3 de Diciembre 1997).
  • Costa Rica, Asamblea Legislativa. Ley Orgánica del Ambiente, No. 7554. La Gaceta, Diario Oficial, No. 215 (13 de Noviembre 1995).
  • Costa Rica, Ministerio de Ambiente y Energía (MINAE). 1997. Ley de Conservación de la Vida Silvestre y su Reglamento, No. 7317. 2 da. Edición. EDITEC EDITORES, S.A.. San José, Costa Rica. 134 p.
  • Frost, D.R. 1985. AMPHIBIANS SPECIES OF THE WORLD: A Taxonomic and Geographical Reference. Published as a Joint Venture of Allen Press, Inc. and The Association Systematics Collections Lawrence, Kansas, U.S.A. 732 p..
  • Frost, Darrel R. 2009. database. Amphibian Species of the World: an Online Reference v5.3
  • Gunther, A.C.L. 1900-02. Biologia Centrali-Americana. Reptilia and Batrachia. 1885-1902.
  • Leenders, T. (2001). A Guide to Amphibians And Reptiles of Costa Rica. Zona Tropical, Miami.
  • Orrell T. (custodian) (2015). ITIS Global: The Integrated Taxonomic Information System (version Mar 2015). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
  • Prohl, H. 1997. Los Anfibios de Hitoy Cerere, Costa Rica. Proyecto Namasol, Cooperación Técnica Bilateral Holanda-Costa Rica. San José, Costa Rica. 66 p.
  • Savage, J. M. (2002). The Amphibians and Reptiles of Costa Rica. University of Chicago Press, Chicago and London.
  • Savage, J.M. 1968. The Dendrobatid Frogs of Central America. Copeia (4): 745-776.
  • Savage, J.M. 1973. A Preliminary Handlist of the Herpetofauna of Costa Rica. s.d.. 17 p..
  • Savage, J.M. 2002. The amphibians and reptiles of Costa Rica. The University of Chicago Press. China. 934 p.
  • Savage, J.M. y J. Villa. 1986. Introduction to the Herpetofauna of Costa Rica/Introducción a la Herpetofauna de Costa Rica. Society for the Studies of Amphibians and Reptiles, Athens, 207.
  • Taylor, E.H. 1952. A review of frogs and toads of Costa Rica. Uni. Kansas Sci. Bull. 35: 577-942.
  • Universidad de Costa Rica. s.f.. CATALOGO DE HERPETOLOGIA. Museo de Zoología, Escuela de Biología, Universidad de Costa Rica. s.d..


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