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BiologyFound in deep and shallow parts of rivers, with moderate to swift current usually at depths of 1 to 8 m (Ref. 57765). Adults live essentially in freshwater although some fish frequently occur in estuaries. Males are sexually mature between 9 and 29 years; females between 9 and 34 years (Ref. 57765). Spawn in main river channel over stone-gravel or gravel-sand bottom and with strong current (Ref. 59043).