Comprehensive DescriptionRead full entry
A small toad-like anuran (males 50-70 mm SVL). Head, viewed from above, rounded, broader than long, about two-thirds as broad as body. Snout blunt, truncate. Nostrils superolateral, located at the tip of the snout. Area between nostrils slightly depressed. Eyes large and prominent, anterolateral. Tympanum not visible. Anterior region of the dorsum with two tooth-like glands of moderate size, and several smaller ones evenly distributed. It also bears a markedly enlarged, smooth temporal gland. Belly granular. Fingers free; a pronounced shovel-like tubercle on the base of the first finger (Cochran 1955). A single moderate glandular ridge is present on the posterior surface of forearm; a greatly enlarged tibial gland is also present. The inner metatarsal tubercle is greatly enlarged and shovel-like, modified for burrowing (Savage and Cei 1965).
Coloration: Dorsal surfaces of the head, back and limbs brown, with faint lighter beige areas on head and some faint black markings on back and limbs. Enlarged warts uniform brown, except for some black edging. A distinct yellow streak along sides, running obliquely downward from level of parotoid gland toward groin. Side of the head and upper lip with regular dark blotches. Belly dirty yellow with some brown pigment. Throat of males dark chocolate brown (Savage and Cei 1965).
Tadpoles: At Gosner 37, body globose, slightly flattened below. Eyes and nostrils dorsal. Body dark brown, with several small dark blotches dorsally; venter finely pigmented with black but transparent. Tail musculature and fins heavily pigmented with a sharp contrast between light and dark areas. Dorsal fin higher than ventral. Spiracle sinistral directed back- and upward, vent tube median, located at the lower part of the body; labial papillae well-developed laterally and along lower labium, one row of marginal papilae, not infolded laterally, dorsal gap present; LTRF 2(2)/3(1); upper and lower jaws finely serrated (Savage and Cei 1965). Lynch (1971) presented a diagnosis for the genus based on osteological characters.
The diploid number is equal to 22 (Savage and Cei 1965).