Crocodylus porosus is most commonly found on the coasts of northern Australia, and the islands of New Guinea and Indonesia. It ranges west as far as the shores of Sri Lanka and eastern India, all along the shorelines and rivermouths of southeast Asia to central Vietnam, around Borneo and into the Philippines, and even out to Palau, Vanuatu, and the Solomon Islands. Saltwater Crocodiles are strong swimmers and can be found very far from land.
(Britton, 1995; Lanworn, 1972; Carr, 1972)
Biogeographic Regions: oriental (Native ); australian (Native ); indian ocean (Native ); pacific ocean (Native )
Distribution: Australia, Bangladesh, Brunei, Myanmar (= Burma), Cambodia, China, India (including Andaman Islands, Nicobar Islands), Indonesia (Java, Sulawesi, Komodo), Malaysia, Palau (Caroline Islands) , Papua New Guinea, Philippines, Singapore, Sri Lanka, Solomon Islands, Thailand, Vanuatu (Banks Islands), Nauru?, Micronesia, Vietnam.
Type locality: Unknown; designated as "Ceylon" (=Sri Lanka) by Mertens (1960: 271).
Southeast Asia, Australia, Papua New Guinea, Islands of the West Pacific Ocean
Saltwater crocodiles are the largest reptilian species alive today. Adult males can reach up to sizes of 6 to 7 meters. Females are much smaller and do not generally exceed 3 meters, with 2.5 meters considered large. The head is very large and a pair of ridges run from the eyes along the center of the snout. The scales are oval in shape and the scutes are small compared to other species. Young saltwater crocodiles are pale yellow in color with black stripes and spots on the body and tail. This coloration lasts for several years until the crocodile matures into an adult. The color as an adult is much darker, with lighter tan or gray areas. The ventral surface is white or yellow in color. Stripes are present on the lower sides of the body but do not extend onto the belly. The tail is gray with dark bands. Saltwater crocodiles have a heavy set jaw which contains up to 68, and no less than 64, teeth.
Range mass: 1000 to 1200 kg.
Other Physical Features: ectothermic ; heterothermic ; bilateral symmetry
Sexual Dimorphism: male larger
- Britton, A. 1995. "Species Account: C. Porosus" (On-line). Accessed March 4, 2001 at http://www.flmnh.ufl.edu/cnhc/csp_cpor.htm.
- Kondo, H. 1970. Grolier's Amazing World of Reptiles. New York, NY: Grolier Interprises Inc.
- Lanworn, R. 1972. The Book of Reptiles. New York, NY: The Hamlyn Publishing Group Ltd.
Habitat and Ecology
The Saltwater Crocodile shows a high tolerance for salinity, being found mostly in coastal waters or around rivers. It may also be found in freshwater rivers, billabongs and swamps.
Movement between habitats occurs during the wet season, when juveniles are raised in freshwater rivers. However, these juveniles are usually forced out of these areas, by dominant males who use the freshwater areas for breeding grounds, and into areas of low salinity. Males who are unable to establish a territory in the river system are either killed or forced out into the sea where they move around the coast in search of another river system.
(Britton, 1995; Pope, 1955)
Terrestrial Biomes: savanna or grassland
Aquatic Biomes: rivers and streams; coastal
The Saltwater Crocodile is a predator and has many different types of prey. When young, Crocodylus porosus is restricted to smaller prey like insects, amphibians, crustaceans and small fish and reptiles. When they become an adult, they feed on larger prey such as mud crabs, turtles, snakes, birds, buffalo, wild boar, and monkeys. When the Saltwater Crocodile hunts for food, it usually hides in the water with only the nostrils, eyes, and part of the back exposed. When the prey approaches, it lunges out of the water and attacks, usually killing its prey with a single snap of the jaws. The Saltwater Crocodile then drags the prey under the water where it is more easily consumed.
Known prey organisms
This list may not be complete but is based on published studies.
Life History and Behavior
Status: captivity: 41.7 years.
Status: captivity: 8.8 years.
Lifespan, longevity, and ageing
The Saltwater Crocodile breeds during the wet season which falls between the months of November and March. Despite the fact that the Saltwater Crocodile is normally found in saltwater areas, breeding grounds are established in fresh water. Males mark out their territory and become defensive if another male tries to enter.
Females reach sexual maturity at around 10 to 12 years old. Males, on the other hand, do not reach sexual maturity until the age of 16 years.
The female crocodile normally lays 40 to 60 eggs, but she can lay up to 90 eggs. The eggs are placed in mounded nests made from plant matter and mud and then buried. Since the eggs are laid during the wet season, the nests must be elevated to prevent loss due to floods.
The male does not stay until the eggs are hatched, but the female stays and protects the nest from predators and humans. After incubation for 90 days, the offspring are hatched, although this time varies with nest temperature. Sex determination is directly related to nest temperature. Males are produced around 31.6 degrees Celsius. If this temperature is increased or decreased just a little, females will be produced. The female unearths the eggs when she hears the chirping sounds the offspring make after they hatch. She then assists the offspring into the water by carrying them in her mouth and tends to them until they learn how to swim.
Range number of offspring: 40 to 90.
Range age at sexual or reproductive maturity (female): 10 to 12 years.
Average age at sexual or reproductive maturity (male): 16 years.
Key Reproductive Features: iteroparous ; seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; oviparous
- Britton, A. 1995. "Species Account: C. Porosus" (On-line). Accessed March 4, 2001 at http://www.flmnh.ufl.edu/cnhc/csp_cpor.htm.
Molecular Biology and Genetics
Barcode data: Crocodylus porosus
Below is a sequence of the barcode region Cytochrome oxidase subunit 1 (COI or COX1) from a member of the species.
See the BOLD taxonomy browser for more complete information about this specimen and other sequences.
-- end --
Download FASTA File
Statistics of barcoding coverage: Crocodylus porosus
Public Records: 4
Specimens with Barcodes: 4
Species With Barcodes: 1
IUCN Red List Assessment
Red List Category
Red List Criteria
- Needs updating
- 1994Vulnerable (V)
- 1990Vulnerable (V)
- 1988Endangered (E)
- 1986Endangered (E)
- 1982Endangered (E)
Although the population of saltwater crocodiles is not stable everywhere, it is in no immediate danger. However, in some countries where the crocodile once thrived, it is now rare or extirpated. Habitat loss associated with coastal development and intensive hunting for hides has drastically reduced populations throughout much of the range. In Sri Lanka and Thailand, habitat destruction is so rapid that the saltwater crocodile has been virtually unseen, with only two saltwater crocodiles being sighted in 1999. In southern Vietnam, where the species once thrived by the thousands, there are but an estimated 100 crocodiles alive in the wild. This is due to the rapid degradation of habitat and the poaching of the animal for leather products. The global population will not be stable until all the countries which have habitats that support the saltwater crocodile have laws that prevent poaching, and programs that create reserves.
A number of such programs have been begun to ensure that C. porosus will not become extinct. In India, a restocking program was introduced in Bhitarkinaka National Park. More than 1,400 saltwater crocodiles were released, with approximately 580 surviving. The population has now become moderately stable at around 1,000 total crocodiles in India. In Burma, crocodile farms are controlling the breeding and conservation of crocodiles. The Australian management program is the world's leader in conservation of the saltwater crocodile. This program focuses its attention on educating the public on precautions to take if they encounter a crocodile, thus discourage unnecessary killing. Crocodile farms were opened to maintain a breeding population, and national sanctuaries have been established, ensuring an undisturbed habitat. Yearly population counts are conducted, monitoring the number of saltwater crocodiles in Australia, making sure that the population does not become dangerously low. In New Papua Guinea, programs that ensure an undisturbed habitat stabilize the population. The Papua New Guinean management system involves a combination of wild cropping, egg and hatchling harvest, and ranching. (Britton, 1995; Carr, 1972)
The IUCN rates the species as a whole as "Low Risk." The U.S. Fish and Wildlife Service rates the Australian population of this species as "Threatened," does not rate the population in Papua New Guinea, and rates the populations in other countries as "Endangered." Saltwater crocodiles from Australia, Indonesia, and Papua New Guinea are included in Appendix II of the CITES treaty, which limits international trade. Members of the species from all other countries are listed in Appendix I, which means they may not be traded internationally.
US Federal List: endangered; threatened
CITES: appendix i; appendix ii
IUCN Red List of Threatened Species: lower risk - least concern
Date Listed: 12/18/1979
Lead Region: Foreign (Region 10)
Where Listed: except Australia & Papua New Guinea
Date Listed: 06/24/1996
Lead Region: Foreign (Region 10)
Where Listed: Australia
Population location: Australia
Listing status: T
Population location: Entire, except Papua New Guinea and Australia
Listing status: E
For most current information and documents related to the conservation status and management of Crocodylus porosus , see its USFWS Species Profile
Relevance to Humans and Ecosystems
The Saltwater Crocodile can be a very dangerous animal to encounter, and humans are attacked and killed by this species every year. Many of these attacks could be prevented by increased awareness and education.
The hide of the Saltwater crocodile is considered very valuable. Many people will pay large amounts of money to have crocodilian products, and Saltwater Crocodile leather products are the most prized. Farms are run for this specific purpose. The crocodile is raised until it is ready to be skinned for leather products. This is a contrversial topic as many people do not find it fit to kill the crocodiles to obtain a small amount of the hide, while the rest of the crocodile is thrown aside.
The saltwater crocodile (Crocodylus porosus), also known as saltie, estuarine or Indo-Pacific crocodile, is the largest of all living reptiles, as well as the largest terrestrial and riparian predator in the world. The males of this species can reach sizes up to 6.7 m (22 ft) and weigh up to 2,000 kg (4,400 lb). However, an adult male saltwater crocodile is generally between 4.3 and 5.2 m (14 and 17 ft) in length and weighs 400 to 1,000 kg (880–2,200 lb), rarely growing larger. Females are much smaller and often do not surpass 3 m (9.8 ft). As the name implies, this species of crocodile can live in salt water, but usually resides in mangrove swamps, estuaries, deltas, lagoons, and lower stretches of rivers. They have the broadest distribution of any modern crocodile, ranging from the eastern coast of India, throughout most of Southeast Asia, stretching south to northern Australia, and historically ranging as far west as just beyond the eastern coast of Africa and as far east as waters off the coast of Japan.
The saltwater crocodile is a formidable and opportunistic hypercarnivorous "apex" ambush predator capable of taking almost any animal that enters its territory, including fish, crustaceans, reptiles, birds and mammals, including other predators. Due to their size and distribution, saltwater crocodiles are the most dangerous extant crocodilian to humans.
- 1 Taxonomy and evolution
- 2 Characteristics
- 3 Distribution and habitat
- 4 Biology and behaviour
- 5 Conservation status
- 6 Relationship with humans
- 7 See also
- 8 References
- 9 External links
Taxonomy and evolution
Crocodylus porosus is believed to have a direct link to similar crocodilians that inhabited the shorelines of the supercontinent Gondwana (which included what is now the Australian continent) as long ago as 98 million years and were survivors of the Cretaceous–Paleogene extinction event. Fossils of Isisfordia, discovered in outback western Queensland (once a vast inland sea) though smaller in size, show attributes of direct lineage to Crocodylus porosus, suggesting it occupied a similar habitat, with vertebrae indicating it shared the ability to death roll during feeding. Incomplete fossil records make it difficult to accurately trace the emergence of the species. The genome was fully sequenced in 2007. The earliest fossil evidence of the species dates to around 4.0-4.5 million years ago  and no subspecies are known. Scientists estimate that C. porosus is an ancient species that could have diverged from 12 to 6 million years ago.
The saltwater crocodile has a wide snout compared to most crocodiles. However, it has a longer muzzle than the mugger crocodile; its length is twice its width at the base. The saltwater crocodile has fewer armour plates on its neck than other crocodilians. On this species, a pair of ridges runs from the eyes along the centre of the snout. The scales are oval in shape and the scutes are small compared to other species. The adult saltwater crocodile's broad body contrasts with that of most other lean crocodiles, leading to early unverified assumptions the reptile was an alligator. The head is very large. Skull lengths of more than 75 cm (30 in) have been confirmed for the species, and mandibular lengths have been reported up to 98.3 cm (38.7 in) (female skull lengths of over 50 cm (20 in) are exceptional). The teeth are also long, with the largest teeth (the fourth tooth from the front on the lower jaw) having been measured to 9 cm (3.5 in) in length. If detached from the body, the head of a very large male crocodile can reportedly weigh over 200 kg (440 lb) alone.
Young saltwater crocodiles are pale yellow in colour with black stripes and spots on their bodies and tails. This colouration lasts for several years until the crocodiles mature into adults. The colour as an adult is much darker greenish-drab, with a few lighter tan or grey areas sometimes apparent. Several colour variations are known and some adults may retain fairly pale skin, whereas others may be so dark as to appear blackish. The ventral surface is white or yellow in colour on saltwater crocodiles of all ages. Stripes are present on the lower sides of their bodies, but do not extend onto their bellies. Their tails are grey with dark bands.
Newly hatched saltwater crocodiles measure about 25 to 30 cm (9.8–11.8 in) long and weigh an average of 70 g (2.5 oz). By their second year, young crocodiles grow to 1 m (3.3 ft) long and weigh 2.5 kg (5.5 lb). Males reach sexual maturity around 3.3 m (11 ft) at around 16 years of age, while females reach sexual maturity at 2.1 m (6.9 ft) and 12–14 years of age. An adult male saltwater crocodile is normally 4.3 to 5.2 m (14–17 ft) long, weighing 400 to 1,000 kg (880–2,200 lb). However, large, mature males can exceed 6 m (20 ft) in length and weigh more than 1,000 kg (2,200 lb). The largest confirmed saltwater crocodile on record was 6.3 m (20.7 ft) long, and weighed over 1,360 kg (3,000 lbs). Due to extensive poaching during the 20th century, such individuals are extremely rare today, as it takes a long time for the crocodiles to attain those sizes. Also, a possible earlier presence of particular genes may have led to such large-sized saltwater crocodiles, genes that were ultimately lost from the overall gene pool due to trophy hunting. However, with recent restoration of saltwater crocodile habitat and reduced poaching, the number of large crocodiles is increasing, especially in Odisha. Recently, Guinness has accepted a claim of a 7.1-metre (23 ft), 2,000-kg (4,400-lb) male saltwater crocodile living within Bhitarkanika Park in Odisha. However, due to the difficulty of trapping and measuring a very large living crocodile, the accuracy of these dimensions has yet to be verified. Currently, four other crocodiles over 6 m (20 ft) are claimed to live in the Bhitarkanika Park. In the future, if conservation efforts pay off, these large individuals could be more common. This species is the only extant crocodilian to regularly reach or exceed 5.2 m (17 ft).
The main determinant of weight is length. The weight of a crocodile increases steeply as length increases. A 10-cm increase in a 1-m-long crocodile would result in a 900-g increase in weight, but a 34-kg gain in the case of a 5-m-long individual. This explains why individuals at 6 m (20 ft) weigh more than twice that of individuals at 5 m (16 ft). Weight can also vary enormously based on condition and age; older males tend to outweigh younger ones, since they maintain prime territories with access to better, more abundant prey. For example, crocodiles at 4.8 m (16 ft) long usually range in weight from 422 to 950 kg (930 to 2,094 lb). On average, though, these 4.8-m individuals would weigh around 520 kg (1,150 lb), and at 5.5 to 5.8 m (18–19 ft) would weigh about 1,000 kg (2,200 lb). This species has the greatest sexual dimorphism of any modern crocodilian, with the females being much smaller than males. Typical female body lengths range from 2.3 to 3.5 m (7.5 to 11.5 ft). Females at 3 m (9.8 ft) in length, such as Connie and Cookie, kept at the Australia Zoo, weigh 150 kg (330 lb). A female of a median length of 2.7 m (8.9 ft) will weigh about 80 to 100 kg (180 to 220 lb). The largest female on record measured about 4.2 m (14 ft).
Distribution and habitat
The saltwater crocodile is one of the three crocodilians found in India, the other two being the more widespread, smaller mugger crocodile and the narrow-snouted, fish-eating gharial. Apart from the eastern coast of India, the saltwater crocodile is extremely rare on the Indian subcontinent. A huge population, consisting of many large adults, several over 6 m (20 ft), including a 7-m (23-ft) male, is present within the Bhitarkanika Wildlife Sanctuary of Odisha and they are known to be present in smaller numbers throughout the Indian and Bangladeshi portions of the Sundarbans. Populations are also present within the mangrove forests and other coastal areas of the Andaman and Nicobar Islands in India. Saltwater crocodiles were once present throughout most of the island of Sri Lanka, but remain mostly within protected areas such as Yala National Park, which also has a large population of mugger crocodiles.
In northern Australia (which includes the northernmost parts of the Northern Territory, Western Australia, and Queensland), the saltwater crocodile is thriving, particularly in the multiple river systems near Darwin (such as the Adelaide, Mary, and Daly Rivers, along with their adjacent billabongs and estuaries), where large individuals of more than 5 m (16 ft) in length are not uncommon. The saltwater crocodile population in Australia is estimated at 100,000 to 200,000 adults. In Australia, the species coexists with the smaller, narrow-snouted Johnston's or freshwater crocodile. Their range extends from Broome in Western Australia through the entire Northern Territory coast all the way south to Rockhampton in Queensland. The Alligator Rivers in the Arnhem Land region are misnamed due to the resemblance of the saltwater crocodile to alligators as compared to freshwater crocodiles, which also inhabit the Northern Territory. In New Guinea, they are also common, existing within the coastal reaches of virtually every river system in the country, such as the Fly River, along with all estuaries and mangroves, where they overlap in range with the rarer, less aggressive New Guinea crocodile. They are also present in varying numbers throughout the Bismarck Archipelago, the Kai Islands, the Aru Islands, the Maluku Islands and many other islands within the region, including Timor, and most islands within the Torres Strait.
The saltwater crocodile was historically known to be widespread throughout Southeast Asia, but is now extinct throughout much of this range. This species has not been reported in the wild for decades in most of Indochina and is extinct in Thailand, Laos, Vietnam, and possibly Cambodia. The status of this species is critical within much of Myanmar, but a stable population of many large adults is present in the Irrawaddy Delta. Probably, the only country in Indochina still harbouring wild populations of this species is Myanmar. Although saltwater crocodiles were once very common in the Mekong Delta (from where they disappeared in the 1980s) and other river systems, the future of this species in Indochina is now looking grim. However, it is also the least likely of crocodilians to become globally extinct due to its wide distribution and almost precolonial population sizes in Northern Australia and New Guinea.
The saltwater crocodile has been long extinct in China, where it inhabited the southern coastal areas from Fujian province in the north to the border of Vietnam. References to large crocodiles that preyed on both humans and livestock appeared during the Han and Song Dynasties, where it occurred in the lower Pearl River near present day Hong Kong and Macau, the Han River, the Min River in the north, portions of coastal Guangxi province and Hainan Island. The presence of crocodiles in Fujian province represent the northernmost distribution of the species.
The population is sporadic in Indonesia and Malaysia, with some areas harbouring large populations (Borneo and Sumatra, for example) and others with very small, at-risk populations (e.g., Peninsular Malaysia). Despite the close proximity to the crocodile hotbed of northern Australia, crocodiles no longer exist in Bali. This species is also reportedly extinct on Lombok, Komodo, and most of Java. A small population may remain within Ujung Kulon National Park in western Java. The saltwater crocodile is also present in very limited parts of the South Pacific, with an average population in the Solomon Islands, a very small and soon to be extinct population in Vanuatu (where the population officially stands at only three) and a decent but at-risk population (which may be rebounding) in Palau. They once ranged as far west as the east coast of Africa to the Seychelles Islands. These crocodiles were once believed to be a population of Nile crocodiles, but they were later proven to be C. porosus.
Because of its tendency to travel very long distances at sea, individual saltwater crocodiles have been known to occasionally appear in areas far away from their general range. Vagrant individuals have historically been reported on New Caledonia, Iwo Jima, Fiji, and even in the relatively frigid Sea of Japan (thousands of miles from their native territory.) In late 2008-early 2009, a handful of wild saltwater crocodiles were verified to be living within the river systems of Fraser Island, hundreds of kilometres from, and in much cooler water than, their normal Queensland range. These crocodiles did indeed migrate south to the island from northern Queensland during the warmer wet season and presumably returned to the north upon the seasonal temperature drop. Despite the surprise and shock within the Fraser Island public, this is apparently not new behaviour, and in the distant past, wild crocodiles had been reported occasionally appearing as far south as Brisbane during the warmer wet season.
Saltwater crocodiles generally spend the tropical wet season in freshwater swamps and rivers, moving downstream to estuaries in the dry season, and sometimes travelling far out to sea. Crocodiles compete fiercely with each other for territory, with dominant males in particular occupying the most eligible stretches of freshwater creeks and streams. Junior crocodiles are thus forced into the more marginal river systems and sometimes into the ocean. This explains the large distribution of the animal (ranging from the east coast of India to northern Australia), as well as its being found in the odd places on occasion (such as the Sea of Japan). Like all crocodiles, they can survive for prolonged periods only in warm temperatures, and crocodiles seasonally vacate parts of Australia if cold spells hit.
Biology and behaviour
The primary behaviour to distinguish the saltwater crocodile from other crocodiles is its tendency to occupy salt water. Though other crocodiles also have salt glands that enable them to survive in saltwater, a trait which alligators do not possess, most other species do not venture out to sea except during extreme conditions. The only other species to display regular seagoing behaviour is the American crocodile, but the American version is still not considered to be as marine-prone as the saltwater crocodile. As its alternate name "sea-going crocodile" implies, this species travels between areas separated by sea, or simply relies on the relative ease of travelling through water in order to circumvent long distances on the same land mass, such as Australia. In a similar fashion to migratory birds using thermal columns, saltwater crocodiles use ocean currents to travel long distances. In a study, 20 crocodiles were tagged with satellite transmitters; 8 of these crocodiles ventured out into open ocean, in which one of them travelled 590 km (370 mi) in 25 days. Another specimen, a 4.84-m-long male, travelled 411 km (255 mi) in 20 days. Without having to move around much, sometimes simply by floating, the current-riding behaviour allows for the conservation of energy. They will even interrupt their travels, residing in sheltered bays for a few days, when the current is against the desired direction of travel, until the current changes direction. Crocodiles also travel up and down in river systems, periodically.
While most crocodilians are social animals sharing basking spots and food, saltwater crocodiles are more territorial and are less tolerant of their own kind; adult males will share territory with females, but drive off rival males. Saltwater crocodiles mate in the wet season, laying eggs in a nest consisting of a mound of mud and vegetation. The female guards the nest and hatchlings from predators.
Generally very lethargic, a trait which helps it survive months at a time without food, the saltwater crocodile will usually loiter in the water or bask in the sun during much of the day, preferring to hunt at night. A study of seasonal saltwater crocodile behaviour in Australia indicated that they are more active and more likely to spend time in the water during the Australian summer; conversely, they are less active and spend relatively more time basking in the sun during the winter. Saltwater crocodiles, however, are among the most active of all crocodilians, spending more time cruising and active, especially in water. They are much less terrestrial than most species of crocodiles, spending less time on land except for basking. At times, they tend to spend weeks at sea in search of land and in some cases, barnacles have been observed growing on crocodile scales, indicative of the long periods they spend at sea.
Despite their relative lethargy, saltwater crocodiles are agile predators and display surprising agility and speed when necessary, usually during strikes at prey. They are capable of explosive bursts of speed when launching an attack from the water. They can also swim at 15 to 18 mph (24 to 29 km/h) in short bursts, around three times as fast as the fastest human swimmers, but when cruising, they usually go at 2 to 3 mph (3.2 to 4.8 km/h). However, stories of crocodiles being faster than a race horse for short distances across land are little more than urban legend. At the water's edge, however, where they can combine propulsion from both feet and tail, their speed can be explosive.
While crocodilian brains are much smaller than those of mammals (as low as 0.05% of body weight in the saltwater crocodile), saltwater crocodiles are capable of learning difficult tasks with very little conditioning, learning to track the migratory route of their prey as the seasons change, and may possess a deeper language ability than currently accepted.
Hunting and diet
Like most crocodilians, saltwater crocodiles are not fastidious in their choice of food, and readily vary their prey selection according to availability, nor are they voracious, as they are able to survive on relatively little food for a prolonged period. Because of their size and distribution, saltwater crocodiles hunt the broadest range of prey species of any modern crocodilian. Hatchlings are restricted to feeding on smaller animals, such as small fish, frogs, insects and small aquatic invertebrates. In addition to these prey, juveniles also take a variety of freshwater and saltwater fish, various amphibians, crustaceans, molluscs, such as large gastropods and cephalopods, birds, small to medium-sized mammals, and other reptiles, such as snakes and lizards. The larger the animal grows, the greater the variety of its diet, although relatively small prey are taken throughout its lifetime. Among crustacean prey, large mud crabs of the genus Scylla are frequently consumed, especially in mangrove habitats. Ground-living birds, such as the emu and different kinds of water birds, are the most commonly preyed upon birds, due to the increased chance of encounter. Even swift-flying birds and bats may be snatched if close to the surface of water, as well as wading birds while these are patrolling the shore looking for food. Mammalian prey of juveniles and subadults are usually as large as the smaller species of ungulates, such as the greater mouse-deer and hog deer. Various mammalian species including, crab-eating macaques, gibbons, porcupines, wallabies, mongoose, civets, flying foxes, hares, rodents, badgers, otters, fishing cats and chevrotains are readily taken when encountered. Large crocodiles, even the oldest males, do not ignore small species when the opportunity arises. However, larger animals are seldom attacked.
Animals taken by adult crocodiles include deer (such as sambar, muntjac and chital), wild boar, tapirs, monkeys, kangaroos, dingos, dholes, jackals, orangutans, turtles, pythons, monitor lizards, Asian antelopes, such as serow and nilgai, and large bovines, such as banteng, water buffalo, and gaur. As a seagoing species, the saltwater crocodile also preys on a variety of saltwater bony fish and other marine animals, including sea snakes, sea turtles (which are usually taken during mating season when the turtles are closer to shore), sea birds, dugongs, rays, and bull sharks Any type of domestic livestock, such as chicken, sheep, pigs, horses and cattle, and domesticated animals/pets may be eaten. Saltwater crocodiles are dominant over other crocodilians, regularly out-competing and occasionally killing and eating other species, as has been recorded largely with freshwater crocodiles in Australia.
Other large predators within their range, such as sun bears, tigers and leopards, are occasionally taken by adult crocodiles. Water buffalo and gaur, which may weigh over a ton, are considered conventionally as the largest prey taken by male crocodiles. Perhaps the only non-marine animal in this species' range that it has not been known to prey on is the adult Asian elephant. It is an extremely powerful animal; in one case, a one-tonne Suffolk stallion known to haul over two tonnes was pulled into water to its demise by a large male saltwater crocodile. In 2011, the first confirmed report of predation of an adult tiger was reported.
As with other crocodilians, their sharp, peg-like teeth are well-suited to seize and tightly grip prey, but not designed to shear flesh. Small prey is simply swallowed whole, while larger animals are forcibly dragged into deep water and drowned. Large prey is then torn into manageable pieces by "death rolling" (the spinning of the crocodile to twist off hunks of meat) or by sudden jerks of the head.
Saltwater crocodiles have the strongest bite of any animal today and a large saltwater crocodile can crush a full-grown bovid's skull between its jaws. A 5.2 m (17 ft)-long saltwater crocodile has been confirmed as having the highest bite force ever recorded for an animal in a laboratory setting, with a bite force value of 36,000 N (3,700 kgf) (surpassing the previous record of 20,840 N (2,125 kgf) made by a 3.9 m (13 ft)-long American alligator). Based on the regression of mean body mass and mean bite force, the bite forces of multiple crocodile species, 6.7 m (22 ft) individuals were estimated at 6,187 kgf (60,670 N) to 7,736 kgf (75,860 N). The study, led by Dr. Gregory M. Erickson, also shed light to the larger, extinct species of crocodilians. Since crocodile anatomy has changed only slightly for the last 80 million years, current data on modern crocodilians can be used to estimate the bite force of extinct species. An 11 to 12 m (36–39 ft) long Deinosuchus would apply a force of 23,100 kgf (227,000 N), twice that of the latest, higher bite force estimations of Tyrannosaurus rex. The extraordinary bite of crocodilians is a result of their anatomy. The space for the jaw muscle in the skull is very large, which is easily visible from the outside as a bulge at each side. The nature of the muscle is extremely stiff, almost as hard as bone to the touch, such that it can appear to be the continuum of the skull. Another trait is that most of the muscle in a crocodile's jaw is arranged for clamping down. Despite the strong muscles to close the jaw, crocodiles have extremely small and weak muscles to open the jaw. The jaws of a crocodile can be securely shut with several layers of duct tape.
Saltwater crocodiles mate in the wet season, when water levels are at their highest. In Australia, the male and female engage in courtship in September and October, and the female lays eggs between November and March. The female selects the nesting site, and both parents will defend the nesting territory, which is typically a stretch of shore along tidal rivers or freshwater areas. The nest is a mound of mud and vegetation, usually measuring 175 cm (69 in) long and 53 cm (21 in) high. The female typically lays from 40 to 60 eggs, but some clutches have included up to 90. The eggs measure on average 8 by 5 cm (3.1 by 2.0 in) and weigh 113 g (4.0 oz) on average. These are relatively small, as the average female saltwater crocodile weighs around five times as much as a freshwater crocodile, but lays eggs that are only about 20% larger in measurement and 40% heavier than those of the smaller species. Although the female guards the nest for 80 to 98 days, the loss of eggs is often high due to flooding and occasionally to predation. As in all crocodilians, the sex of the hatchlings is determined by temperature, with relatively low temperatures producing mainly females, and high temperatures producing mainly males. In Australia, goannas commonly eat freshwater crocodile eggs (feeding on up to 95% of clutch if discovered), but are relatively unlikely to eat saltwater crocodile eggs due to the vigilance of the imposing mother.
As in all crocodilian species, the female saltwater crocodile exhibits a remarkable level of maternal care for a reptile. She excavates the nest in response to "yelping" calls from the hatchlings, and even gently rolls eggs in her mouth to assist hatching. The female will then carry the hatchlings to water in her mouth (as Nile crocodile and American alligator females have been observed doing when their eggs hatch) and remains with the young for several months. Despite her diligence, losses of baby crocodiles are heavy due to various predators and unrelated crocodiles of their own species. Only approximately 1% of the hatchlings will survive to adulthood. The young naturally start to disperse after around 8 months, and start to exhibit territorial behaviour at around 2.5-year-old. However, even females will not reach proper sexual maturity for another 10 years. Saltwater crocodiles that survive to adulthood can attain a very long lifespan, with a life expectancy upwards of 70 years, and some individuals may exceed 100 years. While adults have no natural predators aside from humans, baby saltwater crocodiles may fall prey to monitor lizards, predatory fish, various aquatic and raptorial birds, larger crocodiles, and many other predators. Juveniles may also fall prey to tigers and leopards in certain parts of their range, although encounters between these predators are rare and cats are likely to avoid areas with saltwater crocodiles.
In addition to being hunted for its meat and eggs, the saltwater crocodile has the most commercially valuable skin of any crocodilian; and unregulated hunting during the 20th century caused a dramatic decline in the species throughout its range, with the population in northern Australia reduced 95% by 1971. The species currently has full legal protection in all Australian states and territories where it is found – Western Australia (since 1970), Northern Territory (since 1971) and Queensland (since 1974). Illegal hunting still persists in some areas, with protection in some countries being grossly ineffective, and trade is often difficult to monitor and control over such a vast range. Despite this, the species has made a dramatic recovery in recent decades. Because of its resurgence, the species is considered of minimal concern for extinction. Currently, the species is listed in CITES as follows:
- Appendix I (prohibiting all commercial trade in the species or its byproducts): All wild populations except for those of Australia, Indonesia and Papua New Guinea
- Appendix II (commercial trade allowed with export permit; import permits may or may not be required depending on the laws of the importing country): Australia, Indonesia and Papua New Guinea wild populations, plus all worldwide populations bred in captivity for commercial purposes
Habitat loss continues to be a major problem. In northern Australia, much of the nesting habitat of the saltwater crocodile is susceptible to trampling by feral water buffalo, although buffalo eradication programs have now reduced this problem considerably. Even where large areas of suitable habitat remain, subtle habitat alterations can be a problem, such as in the Andaman Islands, where freshwater areas, used for nesting, are being increasingly converted to human agriculture. After the commercial value of crocodile skins waned, perhaps the greatest immediate challenge to implementing conservation efforts has been the occasional danger the species can be to humans and the resulting negative view of the crocodile.
Relationship with humans
Attacks on humans
Saltwater crocodiles have a strong tendency to treat humans in their territory as prey, and have a long history of attacking humans who unknowingly stray into their territory. As a result of their power, intimidating size and speed, survival of a direct predatory attack is unlikely if the crocodile is able to make direct contact. In distinct contrast to the American policy of encouraging a certain degree of habitat coexistence with alligators, the only recommended policy for dealing with saltwater crocodiles is to avoid their territory whenever possible, as they tend to be highly aggressive when encroached upon.
Exact data on attacks are limited outside Australia, where one or two fatal attacks are reported per year. The low level of attacks may be due to extensive efforts by wildlife officials in Australia to post crocodile warning signs at numerous at-risk billabongs, rivers, lakes and beaches. In the large Aboriginal community of Arnhem Land, attacks frequently go unreported. Also, recent, less-publicised attacks have been reported in Borneo, Sumatra, Eastern India (Andaman Islands), and Burma.
Many attacks in areas outside Australia are believed to go unreported, with one study positing up to 20 to 30 attacks occur every year. Some attacks appear to be territorial rather than predatory in nature, with crocodiles over two years in age often attacking anything that comes into their area (including boats). Humans can usually escape alive from such encounters, which comprise about half of all attacks. Non-fatal attacks usually involve crocodiles of 3 m (9.8 ft) or less in length. Fatal attacks, most likely predatory in nature, commonly involve larger crocodiles with an average estimated size of 4.3 m (14 ft). Under normal circumstances, Nile crocodiles are believed to be responsible for a considerably greater number of fatal attacks on humans than saltwater crocodiles. This is most likely because of the plethora of people in Africa who rely on riparian areas for their livelihood, which is less prevalent in most of Asia and certainly less so in Australia.
During the Japanese retreat in the Battle of Ramree Island on 19 February 1945, saltwater crocodiles may have been responsible for the deaths of over 400 Japanese soldiers. British soldiers encircled the swampland through which the Japanese were retreating, condemning the Japanese to a night in the mangroves, which were home to thousands of saltwater crocodiles. Many Japanese soldiers did not survive this night, but their death mainly as a result of crocodile attacks has been doubted. Another notorious crocodile attack was in 1985, on ecofeminist Val Plumwood, who survived the attack.
According to Wondjina, the mythology of Indigenous Australians, the saltwater crocodile was banished from the fresh water for becoming full of bad spirits and growing too large, unlike the freshwater crocodile, which was somewhat revered. As such, Aboriginal rock art depicting the saltwater crocodile is rare, although examples of up to 3,000 years old can be found in caves in Kakadu and Arnhem land, roughly matching the species distribution. The species is frequently depicted in contemporary aboriginal art.
The species is featured on several postage stamps, including an 1894 State of North Borneo 12-cent stamp; a 1948 Australian 2 shilling stamp depicting an aboriginal rock artwork of the species; a 1966 Republic of Indonesia stamp; a 1994 Palau 20-cent stamp; a 1997 Australian 22-cent stamp; and a 2005 1 Malaysian ringgit postage stamp.
The species has featured in contemporary Australian film and television including the Crocodile Dundee series of film and The Crocodile Hunter television series. There are now several saltwater crocodile-themed parks in Australia.
The crocodile is considered to be holy on Timor. According to legend, the island was formed by a giant crocodile.
Examples of large crocodiles
The largest size recorded for a saltwater crocodile is the subject of considerable controversy. The reason behind unverified sizes is either the case of insufficient/inconclusive data or exaggeration from the hunter's point of view. This section is dedicated to examples of the largest saltwater crocodiles recorded by any individual, amateur or professional, with the aim of satisfying the public interest without creating data pollution. Below, in descending order starting from the largest, are some examples of large crocodiles, confirmed or unconfirmed, recorded throughout history.
- James R. Montgomery, who ran a plantation near to the Lower Kinabatangan Segama Wetlands in Borneo from 1926 to 1932, claimed to have netted, killed, and examined numerous crocodiles well over 6.1 m (20 ft) there, including a specimen he claims measured 10 m (33 ft). However, no one scientifically confirmed any of Montgomery's specimens and no voucher specimens are known.
- A crocodile shot in the Bay of Bengal in 1840 was reported at 10 m (33 ft). This specimen is unconfirmed and no voucher remains are known.
- A crocodile shot in Queensland in 1957, nicknamed Kris the croc, was reported to be 8.63 m (28.3 ft) long, but no verified measurements were made and no remains of this crocodile exist. A "replica" of this crocodile has been made as a tourist attraction.
- A crocodile killed in 1823 at Jalajala on the main island of Luzon in the Philippines was reported at 8.2 m (27 ft).
- The skull of crocodile shot in Odisha, India, was claimed to measure 7.6-metre (25 ft) in life, but when given scholarly examination, was thought to have come from a crocodile of a length no greater than 7 m (23 ft).
- A reported 7.6-m crocodile was killed in the Hooghly River in the Alipore District of Calcutta. However, examinations of the animal's skull actually indicated it ranged from 6.0 to 6.7 m (19.7–22.0 ft).
- In 2006, Guinness accepted a 7.1-metre (23 ft), 2,000-kg (4,400-lb) male saltwater crocodile living within Bhitarkanika Park in Odisha.
- The record size for a crocodile from Papua New Guinea to be considered authentic by Guinness was a 6.32 m (20.7 ft) specimen shot in May 1966 along the northeastern coast. This specimen had a belly girth of 2.74 m (9.0 ft)
- Another seemingly authentic notable New Guinea giant, which drowned after entanglement in a fisherman's net in 1979, measured 6.2 m (20 ft), with a skull length of 72 cm (28 in).
- In September 2011, a 6.17 metres (20.2 ft) specimen was captured alive in the Philippines, making it one of the largest specimens ever reliably measured snout-to-tail. This specimen, nicknamed Lolong and weighing roughly 1,075 kg (2,370 lb), had a past as a possible man-eater and was being kept alive as an attraction in a local zoo. Lolong died on 10 February 2013 due to stress and infection.
- The official, authentic record length for a crocodile from Australia was 6.15 m (20.2 ft) for a specimen killed in the McArthur River in June 1960.
- Another confirmed Australian giant shot 16 years after that, an old male nicknamed "Big Gator" (despite actually being a crocodile, not an alligator) that had become a habitual predator of local cattle, was found after being shot to have measured 6.1 m (20 ft) and weighed 1,097 kilograms (2,418 lb).
- Crocodile Specialist Group (1996). "Crocodylus porosus". IUCN Red List of Threatened Species. Version 2011.1. International Union for Conservation of Nature. Retrieved 20 August 2011.
- Erickson, GM; Gignac PM; Steppan SJ; Lappin AK; Vliet KA et al. (2012). "Insights into the Ecology and Evolutionary Success of Crocodilians Revealed through Bite-Force and Tooth-Pressure Experimentation". PLoS ONE 7 (3): e31781. doi:10.1371/journal.pone.0031781.
- Guinness Book of World Records. Retrieved on 8 April 2013.
- "Our Animals – Reptiles – Crocodilians – Saltwater Crocodile". Australia Zoo. Retrieved 25 July 2013.
- "Saltwater Crocodile facts". Aquaticcommunity.com. Retrieved 22 June 2013.
In Australia, one or two fatal attacks are reported on average per year.
- "Seven-metre man-eating crocodile shot dead". The Daily Telegraph. 15 January 2007.
- Ross, Charles A; Garnett, Stephen, eds. (1989). Crocodiles and Alligators. Checkmark Books. ISBN 978-0816021741.
- Thomas, Abbie; Willis, Paul (July–September 2006). "The Dinosaur Musterers – Dawn of a crocodilian dynasty". Australian Geographic (83): 52–53. Retrieved 8 October 2014.
- "Missing link crocodile found down under". Science Buzz. Science Museum of Minnesota. 18 June 2006. Retrieved 11 July 2013.
- "Ancestor of all modern crocodilians discovered in outback Queensland". The University of Queensland. 14 June 2006. Retrieved 11 July 2013.
- Yan Li, Xiaobing Wu, Xuefeng Ji, Peng Yan & George Amato (February 2007). "The complete mitochondrial genome of salt-water crocodile (Crocodylus porosus) and phylogeny of crocodilians". Journal of genetics and genomics = Yi chuan xue bao 34 (2): 119–128. doi:10.1016/S1673-8527(07)60013-7. PMID 17469784.
- Johnson, David (4 November 2009). The Geology of Australia. Cambridge University Press. p. 181. ISBN 978-0-521-76741-5. Retrieved 28 July 2013.
- Willis, P. M. A., 1997. Review of fossil crocodilians from Australasia. Aust. J. Zool. 30:287–298
- Molnar, R. E., 1979. Crocodylus porosus from the Pliocene Allingham for- mation of North Queensland. Results of the Ray E. Lemley expeditions, part 5. Memoirs Of The Queensland Museum 19:357–365
- Guggisberg, C.A.W. (1972). Crocodiles: Their Natural History, Folklore, and Conservation. Newton Abbot: David & Charles. p. 195. ISBN 0-7153-5272-5.
- Kondo, H. (1970). Grolier's Amazing World of Reptiles. New York, NY: Grolier Interprises Inc.
- Britton, Adam. "Crocodylus porosus (Schneider, 1801)". The Crocodilian Species List.
- Greer, Allen E. (1974). "On the Maximum Total Length of the Salt-Water Crocodile (Crocodylus porosus)". Journal of Herpetology 8 (4): 381–384. doi:10.2307/1562913. JSTOR 1562913.
- Naish, Darren (30 October 2008). "The world's largest modern crocodilian skull – Tetrapod Zoology". Scienceblogs.com. Retrieved 25 July 2013.
- "Australian crocodile Elvis sinks teeth into lawnmower". CNN.com. Retrieved 25 July 2013.
- "Australian Saltwater Crocodile (Estuarine Crocodile) – Crocodylus porosus". Angelfire.com. Retrieved 25 July 2013.
- Wood, Gerald (1983). The Guinness Book of Animal Facts and Feats. p. 256. ISBN 978-0-85112-235-9.
- Lanworn, R. (1972). The Book of Reptiles. New York, NY: The Hamlyn Publishing Group Ltd. ISBN 0600312739.
- "Crocodylus porosus". Kingsnake.com. Retrieved 25 July 2013.
- "Information About Crocodiles". Proserpineecotours.com.au. Retrieved 25 July 2013.
- "Insights into Crocodile Lifestyles". newholland.com.au.
- Mishra, Braja Kishore (14 June 2006). "World's Largest Reptile Found in India". ohmynews.com.
- Hiremath, K. G. (2003). Recent advances in environmental science. Discovery Publishing House. ISBN 81-7141-679-9.
- "West Alligator River". Northern Territory Land Information System. Northern Territory Government. Retrieved 25 September 2012.
- "Crocodile kills man in wildlife sanctuary – World news – World environment – msnbc.com". MSNBC. 20 April 2008. Retrieved 18 August 2010.
- "Current Distribution of Crocodylus porosus". Retrieved 6 February 2014.
- Moskvitch, Katia (7 June 2010). "BBC News – Crocodiles 'surf' long distance on ocean currents". Bbc.co.uk. Retrieved 12 June 2013.
- Grigg, G. C.; Seebacherd, F.; Beard, L. A.; Morris, D. (1998). "Thermal relations of large crocodiles, Crocodylus porosus, free-ranging in a naturalistic situation". Proceedings of the Royal Society B: Biological Sciences 265 (1407): 1793–1799. doi:10.1098/rspb.1998.0504. PMC 1689355.
- "Lepas anatifera Linnaeus, 1758". WallaWalla. Retrieved 2 December 2011.
- "Big Gecko – Crocodile Management, Research and Filming". Crocodilian.com. Retrieved 25 July 2013.
- "Crocodile Communication: Crocodiles, Caimans, Alligators, Gharials". Flmnh.ufl.edu. 5 March 1996. Retrieved 25 July 2013.
- "Emu (Dromaius Novaehollandiae) – Animals – A–Z Animals – Animal Facts, Information, Pictures, Videos, Resources and Links". A–Z Animals. Retrieved 25 July 2013.
- "Crocodile vs. shark photos". photos1.blogger.com.
- "No Bull: Saltwater Crocodile Eats Shark". UnderwaterTimes.com. 13 August 2007. Retrieved 30 April 2011.
- Nelson, D. (2011). "Fifteen-foot Bengali crocodile claims king of jungle title from tiger". Telegraph 11 August 2011. Retrieved 25 July 2013.
- "Crocodiles Have Strongest Bite Ever Measured, Hands-on Tests Show". News.nationalgeographic.com. 15 March 2012. Retrieved 12 June 2013.
- "Saltwater Crocodiles, Crocodylus porosus ~". Marinebio.org. 14 January 2013. Retrieved 25 July 2013.
- Erickson, Gregory M.; Lappin, A. Kristopher; Vliet, Kent A. (2003). "The ontogeny of bite-force performance in American alligator (Alligator mississippiensis)" (PDF). Journal of Zoology 260 (3): 317–327. doi:10.1017/S0952836903003819.
- "Australian saltwater crocodiles are world's most powerful biters". Sciencedaily.com. 16 March 2012. doi:10.1371/journal.pone.0031781. Retrieved 25 July 2013.
- "Crocodilian Species – Australian Saltwater Crocodile (Crocodylus porosus)". Flmnh.ufl.edu. Retrieved 26 December 2010.
- "UNEP-WCMC – Estuarine Crocodile". Web.archive.org. Archived from the original on 29 February 2008. Retrieved 25 July 2013.
- "Crocodylus porosus – Salt-water Crocodile, Estuarine Crocodile". Environment.gov.au. Retrieved 8 August 2013.
- Caldicott, David G.E. et al. (September 2005). "Crocodile Attack in Australia: An Analysis of Its Incidence and Review of the Pathology and Management of Crocodilian Attacks in General". Wilderness & Environmental Medicine 16 (3): 143–159. doi:10.1580/1080-6032(2005)16[143:CAIAAA]2.0.CO;2. PMID 16209470.
- "Rise in Borneo crocodile attacks may be linked to Malaysia's logging, palm oil, expert says". Associated Press via International Herald Tribune. 25 April 2007. Archived from the original on 11 August 2007. Retrieved 18 August 2010.
- "Woman saves daughter from crocodile". Telegraph. 14 March 2008. Retrieved 25 July 2013.
- "KalingaTimes.com: Two injured in crocodile attack in Orissa". Web.archive.org. 7 June 2008. Archived from the original on 7 June 2008. Retrieved 25 July 2013.
- "Croc kills woman 4 years after her sister's death – TODAY.com". Today.msnbc.msn.com. 5 October 2010. Retrieved 25 July 2013.
- "Crocodile kills man accused of illegal logging in Myanmar wildlife sanctuary". Associated Press via International Herald Tribune. 20 April 2008. Archived from the original on 17 May 2008. Retrieved 18 August 2010.
- "Crocodile Specialist Group – Crocodilian Attacks". Web.archive.org. Archived from the original on 3 May 2009. Retrieved 25 July 2013.
- Platt, S. G., W. K. Ko, M. Kalyar Myo, L. L. Khaing, and T. Rainwater. Man eating by estuarine crocodiles: the Ramree Island massacre revisited. Herp Bull 2001. 75:15–18
- "National Museum of Australia – Val Plumwood canoe". Nma.gov.au. Retrieved 25 July 2013.
- "Val Plumwood Prey to a Crocodile". Aislingmagazine.com. Retrieved 25 July 2013.
- "Historical Feature – Aboriginal Dreamtime Legends". Australianstamp.com. 5 December 2007. Retrieved 8 August 2013.
- "Monster croc? Welcome to my nightmare". The Sunday Mail (news.com.au). 28 November 2010. Retrieved 2 April 2011.
- Reid, Robert (28 November 2008). "Death of a monster". The Australian. Retrieved 2 April 2011.
- "QLD – NORMANTON – Home of largest Crocodile ever shot!". Give An Aussie A Go. Retrieved 25 July 2013.
- "Krys the crocodile, Normanton, Queensland :: Backyard". abc.net.au. 6 January 2004.
- "Saltwater Crocodiles, Saltwater Crocodile Pictures, Saltwater Crocodile Facts – National Geographic". Animals.nationalgeographic.com. Retrieved 25 July 2013.
- "Guinness: India Park Home to World's Largest Crocodile". UnderwaterTimes.com. Retrieved 30 April 2011.
- "Crocodilian Biology Database. FAQ". Crocodilian.com. Retrieved 24 May 2012.
- ""Lolong" holds world record as largest croc in the world". Pawb.gov.ph. 17 November 2011. Retrieved 24 May 2012.
- "Croc Blog: Accurate length measurement for Lolong". Crocodilian.blogspot.com. 12 November 2011. Retrieved 25 July 2013.
- "Philippine town claims world's largest crocodile title". The Daily Telegraph (London). 9 November 2011.
- "'Lolong' claims world's largest croc title". Agence France-Presse via abs-cbnnews.com. 11 September 2011.
- "BBC News – Philippine giant croc captured after three-week hunt". Bbc.co.uk. 6 September 2011. Retrieved 25 July 2013.
- "Pictures: Biggest Crocodile Ever Caught?". News.nationalgeographic.com. 6 September 2011. Retrieved 25 July 2013.
- Ortiz, Erik (10 February 2013). "Tears for a croc: Lolong, the world's largest crocodile in captivity, dies in the Philippines". New York Daily News. Retrieved 11 February 2013.