HabitSprawling perennial herbs, woody at the base, to 1 m tall, to 1 m in diameter. Stem 7-10 mm in diameter at base, green, minutely puberulent with simple, uniseriate, stiff 1-2-celled trichomes from a unicellular base, mixed with scattered simple uniseriate glandular trichomes with 4-celled heads, glabrescent in age, all trichomes less than 0.5 mm long.
Sympodial StructureSympodial units 2-foliate (rarely 3-foliate); internodes 2-6 cm long.
LeavesLeaves interrupted imparipinnate, 3.5-13 (-16) cm long, 1-6 (-8) cm wide, bright green, minutely pubescent with stiff simple uniseriate trichomes like those of the stems, pubescence more abundant abaxially, especially along the veins; primary leaflets 3-7 pairs, gradually becoming smaller towards the base of the leaf, subopposite to alternate, narrowly elliptic, the base truncate, more or less oblique basiscopically, the margins deeply and irregularly lobed to occasionally almost entire in some leaflets, the apex acute to acuminate; terminal leaflet 1.2-2.5 cm long, 0.4-1.5 cm wide, the petiolule 0.2-0.5 (-1) cm long, the apex acute to acuminate; lateral leaflets 0.7-2.5 cm long, 0.3-1.2 cm wide, the petiolule 0-0.5 (1) cm long, usually decurrent on the rachis basiscopically; secondary leaflets occasionally present in some leaflets, especially acroscopically, 0.3-0.5 cm long, 0.25-0.4 cm wide, sessile and decurrent on the leaflet rachis; tertiary leaflets absent; interjected leaflets (0-) 3-12 (-20), 0.2-0.5 cm long, 0.1-0.4 cm wide, sessile and decurrent on the main leaf rachis, often 2 sets of unpaired interjected leaflets between each set of primary lateral leaflets; petiole 1-4 (-6) cm long; pseudostipules present or absent, if present then present on most nodes, 0.3-0.4 cm long, 0.2-0.3 cm wide.
InflorescencesInflorescences (6-) 12-30 cm long, once branched, usually regularly bifurcate, with 8-30 flowers, ebracteate or bracteate on most nodes from the base, the bracts 0.2-0.6 cm long, 0.1-0.4 cm wide, the largest bract at the bifurcation and first nodes, peduncle 5-15 cm long, minutely pubescent like the stems, with more prominent and numerous glandular trichomes especially at the apex. Pedicels 0.8-1.6 cm long, articulated in the distal half. Buds 1-1.2 cm long, 0.4-0.5 cm wide, narrowly conical, straight or curved at the very tip, with the corolla more than half exserted from the calyx just before anthesis.
FlowersFlowers with the calyx tube minute, ca. 0.05-0.1 cm long, the sinuses often hyaline, the calyx lobes 0.25-0.5 cm long, 0.1-0.15 cm wide, lanceolate or in some populations (Río Fortaleza) the tips rounded, minutely pubescent like the rest of the inflorescence with short, simple uniseriate 1-2-celled white trichomes; corolla 1.8-2.5 cm in diameter, stellate, yellow, the tube 0.2-0.5 cm long, the lobes 1-1.5 cm long, 0.3-0.7 cm wide, uniformly and sparsely pubescent abaxially with 1-celled uniseriate trichomes, these not more abundant along the midveins, the lobe tips elongate and acuminate, reflexed at anthesis; staminal column 1-1.2 cm long, straight or curved, the filaments less than 0.5 mm long, the anthers 0.4-0.6 cm long, the sterile apical appendage 0.15-0.4 cm long; ovary conical to globose, minutely and sparsely pubescent with simple trichomes at the apex; style 0.8-1.2 cm long, ca. 0.5 mm in diameter, sparsely pubescent in the basal half, exserted 0.5-1.7 mm from the staminal column; stigma capitate, green.
FruitsFruit 1-1.4 cm in diameter, globose, 2-locular, green with a dark green or purple stripe from the apex to the base at fruit maturity, sparsely pubescent with weak-walled simple uniseriate trichomes to 0.5 mm long and occasionally also with minute glandular trichomes with 4-celled heads; fruiting pedicels 1-2 cm long, straight or slightly incurved towards the inflorescence axis; calyx lobes in fruit 1.1-1.6 cm long, 0.15-0.2 cm wide, spreading in some populations (Fortaleza) the lobe tips recurved.
SeedsSeeds 2.4-3 mm long, 1.1-1.5 mm wide, 0.5-0.6 mm thick, obovate, pale brown, pubescent with hair-like outgrowths of the lateral testa cell walls, which give a silky appearance to the surface, narrowly winged at the apex and acute at the base.
Catalog Number: US 2591186A
Collection: Smithsonian Institution, National Museum of Natural History, Department of Botany
Verification Degree: Status verified from secondary sources
Preparation: Pressed specimen
Collector(s): E. C. Smith & J. Blas
Year Collected: 1970
Locality: Callejon de Huaylas, Dept. Ancash. Trail to cave across Rio Santa from Mancos., Ancash, Peru, South America
- Isotype: Peralta, I. & Knapp, S. 2005. Syst. Bot. 30: 431.
Life History and Behavior
Description[edit source | edit]
It is a sprawling perennial herb, woody at the base, the herb being up to 1 metre (3.3 ft) or more in diameter and up to 1m tall. Its stem diameter ranges between 7 and 10 millimetres (0.28 and 0.39 in) at the base, not hollow in age, green, minutely puberulent with simple, uniseriate, stiff 1–2-celled trichomes. Sympodial units are 2-foliate (sometimes 3-foliate); internodes between 2 and 6 centimetres (0.79 and 2.4 in). Its leaves are interrupted imparipinnate, bright green, minutely pubescent with stiff simple uniseriate trichomes like those of the stems.
The petiole is between 1 and 4 centimetres (0.39 and 1.6 in); pseudostipules present or absent, if present then present on most nodes. Inﬂorescences range between 12 and 30 centimetres (4.7 and 12 in) in size, with 8–30 ﬂowers, ebracteate or bracteate on most nodes from the base. Peduncle is between 5 and 15 centimetres (2.0 and 5.9 in); pedicels are between 0.8 and 1.6 centimetres (0.31 and 0.63 in), articulated in the upper half. Flowers with the calyx tube are minute, approximately between 0.5 and 1 millimetre (0.020 and 0.039 in). Fruit is between 1 and 1.4 centimetres (0.39 and 0.55 in) in diameter, globose and green when ripe. Seeds are obovate, narrowly winged at the apex and acute at the base, pale brown, pubescent with hair-like outgrowths of the tegument cell radial walls, which give the surface a silky appearance. Chromosome number: n=12 .
Distribution[edit source | edit]
On rocky slopes in the Department of Ancash, Peru at an altitude between 940 and 3,000 metres (3,080 and 9,800 ft).
References[edit source | edit]
Further reading[edit source | edit]
- Stadler, T.; Arunyawat, U.; Stephan, W. (2008). "Population Genetics of Speciation in Two Closely Related Wild Tomatoes (Solanum Section Lycopersicon)". Genetics 178 (1): 339–350. doi:10.1534/genetics.107.081810. ISSN 0016-6731.
- Cortada et al. (2010). "THE RESISTANCE RESPONSE OF SOLANUM HUAYLASENSE ACCESSION LA1358 TO MELOIDOGYNE SPP.". NEMATROPICA 40 (1): 31–40. Retrieved 15 July 2013.
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