The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • 1996. database, The PLANTS Database
  • 1996. database, NODC Taxonomic Code
  • 2000. database, The PLANTS Database
  • 2011. Lamiales of North America Update
  • Flora of China Editorial Committee. 1994. Flora of China (Verbenaceae through Solanaceae). 17: 1–378. In C. Y. Wu, P. H. Raven & D. Y. Hong (eds.) Fl. China. Science Press & Missouri Botanical Garden Press, Beijing & St. Louis.
  • Nasir, E. & S. I. Ali (eds). 1980-2005. Fl. Pakistan Univ. of Karachi, Karachi.
  • Orrell T. (custodian) (2015). ITIS Regional: The Integrated Taxonomic Information System (version Dec 2014). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
  • Species Plantarum 2: 595. 1753. (1 May 1753) (Sp. Pl.)


EOL content is automatically assembled from many different content providers. As a result, from time to time you may find pages on EOL that are confusing.

To request an improvement, please leave a comment on the page. Thank you!