The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • 1996. database, The PLANTS Database
  • 1996. database, NODC Taxonomic Code
  • 2000. database, The PLANTS Database
  • 2007. Flora of the West Indies website
  • Boraginaceae and Hydroleaceae of North America Update
  • Gibson, D. 1970. Boraginaceae. Flora of Guatemala. Fieldiana Botany. Vol. 24, parte IX: 111-167.
  • Jimémez, Q. 1999. Árboles maderables en peligro de extinción en Costa Rica.II edición. Instituto Nacional de Biodiversidad. 163 p.
  • Orrell T. (custodian) (2015). ITIS Regional: The Integrated Taxonomic Information System (version Dec 2014). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
  • Zamora, N. 1989. Flora arborescente de Costa Rica. Editorial Tecnológica de Costa Rica. 262 p.


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