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  <id type="integer">925983</id>
  <canonical-form>Hyaena brunnea</canonical-form>
  <iucn-conservation-status>Near Threatened (NT)</iucn-conservation-status>
  <scientific-name>&lt;i&gt;Hyaena brunnea&lt;/i&gt; Thunberg, 1820</scientific-name>
  <common-names>
    <item>
      <language_label>af</language_label>
      <string>Strandwolf</string>
    </item>
    <item>
      <language_label>br</language_label>
      <string>Bleiz-broc'h gell</string>
    </item>
    <item>
      <language_label>da</language_label>
      <string>Brun hy&#230;ne</string>
    </item>
    <item>
      <language_label>en</language_label>
      <string>Brown hyaena</string>
    </item>
    <item>
      <language_label>en</language_label>
      <string>Brown Hyena</string>
    </item>
    <item>
      <language_label>es</language_label>
      <string>Parahyaena brunnea</string>
    </item>
    <item>
      <language_label>fi</language_label>
      <string>Ruskohyeena</string>
    </item>
    <item>
      <language_label>he</language_label>
      <string>&#1510;&#1489;&#1493;&#1506; &#1495;&#1493;&#1501;</string>
    </item>
    <item>
      <language_label>hu</language_label>
      <string>Barna hi&#233;na</string>
    </item>
    <item>
      <language_label>ja</language_label>
      <string>&#12459;&#12483;&#12471;&#12519;&#12463;&#12495;&#12452;&#12456;&#12490;</string>
    </item>
    <item>
      <language_label>ka</language_label>
      <string>&#4315;&#4308;&#4316;&#4304;&#4318;&#4312;&#4320;&#4308; &#4304;&#4324;&#4311;&#4304;&#4320;&#4312;</string>
    </item>
    <item>
      <language_label>ko</language_label>
      <string>&#44040;&#49353;&#54616;&#51060;&#50640;&#45208;</string>
    </item>
    <item>
      <language_label>nl</language_label>
      <string>Bruine hyena</string>
    </item>
    <item>
      <language_label>no</language_label>
      <string>Brunhyene</string>
    </item>
    <item>
      <language_label>pl</language_label>
      <string>Hiena brunatna</string>
    </item>
    <item>
      <language_label>pt</language_label>
      <string>Hiena-castanha</string>
    </item>
    <item>
      <language_label>sv</language_label>
      <string>Brun hyena</string>
    </item>
    <item>
      <language_label>tr</language_label>
      <string>Kahverengi s&#305;rtlan</string>
    </item>
  </common-names>
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        <full-reference>Nowak, R. 1999. Walker's Mammals of the World. Baltimore: The John Hopkins University Press.</full-reference>
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      <ref>
        <full-reference>Grzimek, B. 1990. Grzimek's Encyclopedia of Mammals. New York: McGraw-Hill Publishing Co..</full-reference>
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        <full-reference>Mills, M. 23 Nov. 1982. Hyaena brunnea. Mammalian Species, No. 194: 1-5.</full-reference>
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        <full-reference>Owens, D., M. Owens. 1996. Social dominance and reproductive patterns in brown hyaenas, Hyaena brunnea, of the central Kalahari desert. Animal Behavior, vol. 51: 535-551.</full-reference>
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        <full-reference>Nowak, R. 1999. Walker's Mammals of the World. Baltimore: The John Hopkins University Press.</full-reference>
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        <full-reference>Grzimek, B. 1990. Grzimek's Encyclopedia of Mammals. New York: McGraw-Hill Publishing Co..</full-reference>
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        <full-reference>Mills, M. 23 Nov. 1982. Hyaena brunnea. Mammalian Species, No. 194: 1-5.</full-reference>
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        <full-reference>Owens, D., M. Owens. 1996. Social dominance and reproductive patterns in brown hyaenas, Hyaena brunnea, of the central Kalahari desert. Animal Behavior, vol. 51: 535-551.</full-reference>
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        <full-reference>Nowak, R. 1999. Walker's Mammals of the World. Baltimore: The John Hopkins University Press.</full-reference>
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        <full-reference>Grzimek, B. 1990. Grzimek's Encyclopedia of Mammals. New York: McGraw-Hill Publishing Co..</full-reference>
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        <full-reference>Mills, M. 23 Nov. 1982. Hyaena brunnea. Mammalian Species, No. 194: 1-5.</full-reference>
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        <full-reference>Owens, D., M. Owens. 1996. Social dominance and reproductive patterns in brown hyaenas, Hyaena brunnea, of the central Kalahari desert. Animal Behavior, vol. 51: 535-551.</full-reference>
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      <ref>
        <full-reference>1. IUCN Red List  (December, 2007) &lt;a href=&quot;http://www.iucnredlist.org&quot;&gt;http://www.iucnredlist.org&lt;/a&gt;</full-reference>
        <id type="integer">686885</id>
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      <ref>
        <full-reference>2. Kingdon, J. (1997) &lt;i&gt;The Kingdon Field Guide to African Mammals&lt;/i&gt;. Academic Press Limited, London.</full-reference>
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        <published type="integer">1</published>
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        <full-reference>3. Stuart, C. and Stuart, T. (1997) &lt;i&gt;Field Guide to the Larger Mammals of Africa&lt;/i&gt;. Struik Publishers, London.</full-reference>
        <id type="integer">691377</id>
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      <ref>
        <full-reference>4. Wiesel, I. (2008) &lt;i&gt;Pers. Comm.&lt;/i&gt;</full-reference>
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      <ref>
        <full-reference>5. Mills, M.G.L. and Bearder, S.K. (2006) Hyena Family. In: Macdonald, D.W. Ed. &lt;i&gt;The Encyclopedia of Mammals&lt;/i&gt;. Oxford University Press, Oxford.</full-reference>
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      <ref>
        <full-reference>6. Mills, G. and Hes, L. (1997) &lt;i&gt;The Complete Book of Southern African Mammals&lt;/i&gt;. Struik Publishers, Cape Town.</full-reference>
        <id type="integer">691380</id>
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      <ref>
        <full-reference>7. IUCN Hyaena Specialist Group  (December, 2007) &lt;a href=&quot;http://www.hyaenidae.org/&quot;&gt;http://www.hyaenidae.org/&lt;/a&gt;</full-reference>
        <id type="integer">691381</id>
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    <refs type="array">
      <ref>
        <full-reference>1. IUCN Red List  (December, 2007) &lt;a href=&quot;http://www.iucnredlist.org&quot;&gt;http://www.iucnredlist.org&lt;/a&gt;</full-reference>
        <id type="integer">686885</id>
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      <ref>
        <full-reference>2. Kingdon, J. (1997) &lt;i&gt;The Kingdon Field Guide to African Mammals&lt;/i&gt;. Academic Press Limited, London.</full-reference>
        <id type="integer">690093</id>
        <published type="integer">1</published>
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      <ref>
        <full-reference>3. Stuart, C. and Stuart, T. (1997) &lt;i&gt;Field Guide to the Larger Mammals of Africa&lt;/i&gt;. Struik Publishers, London.</full-reference>
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        <full-reference>4. Wiesel, I. (2008) &lt;i&gt;Pers. Comm.&lt;/i&gt;</full-reference>
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        <published type="integer">1</published>
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      <ref>
        <full-reference>5. Mills, M.G.L. and Bearder, S.K. (2006) Hyena Family. In: Macdonald, D.W. Ed. &lt;i&gt;The Encyclopedia of Mammals&lt;/i&gt;. Oxford University Press, Oxford.</full-reference>
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      <ref>
        <full-reference>6. Mills, G. and Hes, L. (1997) &lt;i&gt;The Complete Book of Southern African Mammals&lt;/i&gt;. Struik Publishers, Cape Town.</full-reference>
        <id type="integer">691380</id>
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        <user-submitted type="boolean">false</user-submitted>
        <visibility-id type="integer">1</visibility-id>
      </ref>
      <ref>
        <full-reference>7. IUCN Hyaena Specialist Group  (December, 2007) &lt;a href=&quot;http://www.hyaenidae.org/&quot;&gt;http://www.hyaenidae.org/&lt;/a&gt;</full-reference>
        <id type="integer">691381</id>
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      <ref>
        <full-reference>1. IUCN Red List  (December, 2007) &lt;a href=&quot;http://www.iucnredlist.org&quot;&gt;http://www.iucnredlist.org&lt;/a&gt;</full-reference>
        <id type="integer">686885</id>
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      <ref>
        <full-reference>2. Kingdon, J. (1997) &lt;i&gt;The Kingdon Field Guide to African Mammals&lt;/i&gt;. Academic Press Limited, London.</full-reference>
        <id type="integer">690093</id>
        <published type="integer">1</published>
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        <taxon-id type="NilClass">2453462</taxon-id>
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      <ref>
        <full-reference>3. Stuart, C. and Stuart, T. (1997) &lt;i&gt;Field Guide to the Larger Mammals of Africa&lt;/i&gt;. Struik Publishers, London.</full-reference>
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        <full-reference>4. Wiesel, I. (2008) &lt;i&gt;Pers. Comm.&lt;/i&gt;</full-reference>
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      <ref>
        <full-reference>5. Mills, M.G.L. and Bearder, S.K. (2006) Hyena Family. In: Macdonald, D.W. Ed. &lt;i&gt;The Encyclopedia of Mammals&lt;/i&gt;. Oxford University Press, Oxford.</full-reference>
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      <ref>
        <full-reference>6. Mills, G. and Hes, L. (1997) &lt;i&gt;The Complete Book of Southern African Mammals&lt;/i&gt;. Struik Publishers, Cape Town.</full-reference>
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      <ref>
        <full-reference>7. IUCN Hyaena Specialist Group  (December, 2007) &lt;a href=&quot;http://www.hyaenidae.org/&quot;&gt;http://www.hyaenidae.org/&lt;/a&gt;</full-reference>
        <id type="integer">691381</id>
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      <ref>
        <full-reference>1. IUCN Red List  (December, 2007) &lt;a href=&quot;http://www.iucnredlist.org&quot;&gt;http://www.iucnredlist.org&lt;/a&gt;</full-reference>
        <id type="integer">686885</id>
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      <ref>
        <full-reference>2. Kingdon, J. (1997) &lt;i&gt;The Kingdon Field Guide to African Mammals&lt;/i&gt;. Academic Press Limited, London.</full-reference>
        <id type="integer">690093</id>
        <published type="integer">1</published>
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      <ref>
        <full-reference>3. Stuart, C. and Stuart, T. (1997) &lt;i&gt;Field Guide to the Larger Mammals of Africa&lt;/i&gt;. Struik Publishers, London.</full-reference>
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      <ref>
        <full-reference>4. Wiesel, I. (2008) &lt;i&gt;Pers. Comm.&lt;/i&gt;</full-reference>
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        <published type="integer">1</published>
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      <ref>
        <full-reference>5. Mills, M.G.L. and Bearder, S.K. (2006) Hyena Family. In: Macdonald, D.W. Ed. &lt;i&gt;The Encyclopedia of Mammals&lt;/i&gt;. Oxford University Press, Oxford.</full-reference>
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      <ref>
        <full-reference>6. Mills, G. and Hes, L. (1997) &lt;i&gt;The Complete Book of Southern African Mammals&lt;/i&gt;. Struik Publishers, Cape Town.</full-reference>
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      <ref>
        <full-reference>7. IUCN Hyaena Specialist Group  (December, 2007) &lt;a href=&quot;http://www.hyaenidae.org/&quot;&gt;http://www.hyaenidae.org/&lt;/a&gt;</full-reference>
        <id type="integer">691381</id>
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      <ref>
        <full-reference>1. IUCN Red List  (December, 2007) &lt;a href=&quot;http://www.iucnredlist.org&quot;&gt;http://www.iucnredlist.org&lt;/a&gt;</full-reference>
        <id type="integer">686885</id>
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        <full-reference>2. Kingdon, J. (1997) &lt;i&gt;The Kingdon Field Guide to African Mammals&lt;/i&gt;. Academic Press Limited, London.</full-reference>
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        <full-reference>3. Stuart, C. and Stuart, T. (1997) &lt;i&gt;Field Guide to the Larger Mammals of Africa&lt;/i&gt;. Struik Publishers, London.</full-reference>
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        <full-reference>4. Wiesel, I. (2008) &lt;i&gt;Pers. Comm.&lt;/i&gt;</full-reference>
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        <full-reference>5. Mills, M.G.L. and Bearder, S.K. (2006) Hyena Family. In: Macdonald, D.W. Ed. &lt;i&gt;The Encyclopedia of Mammals&lt;/i&gt;. Oxford University Press, Oxford.</full-reference>
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      <ref>
        <full-reference>6. Mills, G. and Hes, L. (1997) &lt;i&gt;The Complete Book of Southern African Mammals&lt;/i&gt;. Struik Publishers, Cape Town.</full-reference>
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        <full-reference>7. IUCN Hyaena Specialist Group  (December, 2007) &lt;a href=&quot;http://www.hyaenidae.org/&quot;&gt;http://www.hyaenidae.org/&lt;/a&gt;</full-reference>
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        <full-reference>1. IUCN Red List  (December, 2007) &lt;a href=&quot;http://www.iucnredlist.org&quot;&gt;http://www.iucnredlist.org&lt;/a&gt;</full-reference>
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      <ref>
        <full-reference>2. Kingdon, J. (1997) &lt;i&gt;The Kingdon Field Guide to African Mammals&lt;/i&gt;. Academic Press Limited, London.</full-reference>
        <id type="integer">690093</id>
        <published type="integer">1</published>
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      <ref>
        <full-reference>3. Stuart, C. and Stuart, T. (1997) &lt;i&gt;Field Guide to the Larger Mammals of Africa&lt;/i&gt;. Struik Publishers, London.</full-reference>
        <id type="integer">691377</id>
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      <ref>
        <full-reference>4. Wiesel, I. (2008) &lt;i&gt;Pers. Comm.&lt;/i&gt;</full-reference>
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      <ref>
        <full-reference>5. Mills, M.G.L. and Bearder, S.K. (2006) Hyena Family. In: Macdonald, D.W. Ed. &lt;i&gt;The Encyclopedia of Mammals&lt;/i&gt;. Oxford University Press, Oxford.</full-reference>
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      <ref>
        <full-reference>6. Mills, G. and Hes, L. (1997) &lt;i&gt;The Complete Book of Southern African Mammals&lt;/i&gt;. Struik Publishers, Cape Town.</full-reference>
        <id type="integer">691380</id>
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      <ref>
        <full-reference>7. IUCN Hyaena Specialist Group  (December, 2007) &lt;a href=&quot;http://www.hyaenidae.org/&quot;&gt;http://www.hyaenidae.org/&lt;/a&gt;</full-reference>
        <id type="integer">691381</id>
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        <full-reference>1. IUCN Red List  (December, 2007) &lt;a href=&quot;http://www.iucnredlist.org&quot;&gt;http://www.iucnredlist.org&lt;/a&gt;</full-reference>
        <id type="integer">686885</id>
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        <visibility-id type="integer">1</visibility-id>
      </ref>
      <ref>
        <full-reference>2. Kingdon, J. (1997) &lt;i&gt;The Kingdon Field Guide to African Mammals&lt;/i&gt;. Academic Press Limited, London.</full-reference>
        <id type="integer">690093</id>
        <published type="integer">1</published>
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      <ref>
        <full-reference>3. Stuart, C. and Stuart, T. (1997) &lt;i&gt;Field Guide to the Larger Mammals of Africa&lt;/i&gt;. Struik Publishers, London.</full-reference>
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        <full-reference>4. Wiesel, I. (2008) &lt;i&gt;Pers. Comm.&lt;/i&gt;</full-reference>
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      <ref>
        <full-reference>5. Mills, M.G.L. and Bearder, S.K. (2006) Hyena Family. In: Macdonald, D.W. Ed. &lt;i&gt;The Encyclopedia of Mammals&lt;/i&gt;. Oxford University Press, Oxford.</full-reference>
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      <ref>
        <full-reference>6. Mills, G. and Hes, L. (1997) &lt;i&gt;The Complete Book of Southern African Mammals&lt;/i&gt;. Struik Publishers, Cape Town.</full-reference>
        <id type="integer">691380</id>
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      <ref>
        <full-reference>7. IUCN Hyaena Specialist Group  (December, 2007) &lt;a href=&quot;http://www.hyaenidae.org/&quot;&gt;http://www.hyaenidae.org/&lt;/a&gt;</full-reference>
        <id type="integer">691381</id>
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        <full-reference>1. IUCN Red List  (December, 2007) &lt;a href=&quot;http://www.iucnredlist.org&quot;&gt;http://www.iucnredlist.org&lt;/a&gt;</full-reference>
        <id type="integer">686885</id>
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      <ref>
        <full-reference>2. Kingdon, J. (1997) &lt;i&gt;The Kingdon Field Guide to African Mammals&lt;/i&gt;. Academic Press Limited, London.</full-reference>
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      <ref>
        <full-reference>3. Stuart, C. and Stuart, T. (1997) &lt;i&gt;Field Guide to the Larger Mammals of Africa&lt;/i&gt;. Struik Publishers, London.</full-reference>
        <id type="integer">691377</id>
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        <full-reference>4. Wiesel, I. (2008) &lt;i&gt;Pers. Comm.&lt;/i&gt;</full-reference>
        <id type="integer">691378</id>
        <published type="integer">1</published>
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      <ref>
        <full-reference>5. Mills, M.G.L. and Bearder, S.K. (2006) Hyena Family. In: Macdonald, D.W. Ed. &lt;i&gt;The Encyclopedia of Mammals&lt;/i&gt;. Oxford University Press, Oxford.</full-reference>
        <id type="integer">691379</id>
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      <ref>
        <full-reference>6. Mills, G. and Hes, L. (1997) &lt;i&gt;The Complete Book of Southern African Mammals&lt;/i&gt;. Struik Publishers, Cape Town.</full-reference>
        <id type="integer">691380</id>
        <published type="integer">1</published>
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        <user-submitted type="boolean">false</user-submitted>
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      <ref>
        <full-reference>7. IUCN Hyaena Specialist Group  (December, 2007) &lt;a href=&quot;http://www.hyaenidae.org/&quot;&gt;http://www.hyaenidae.org/&lt;/a&gt;</full-reference>
        <id type="integer">691381</id>
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        <full-reference>Richard Weigl (2005) Longevity of Mammals in Captivity; from the Living Collections of the World. Kleine Senckenberg-Reihe 48: Stuttgart.</full-reference>
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        <full-reference>Ronald Nowak (1999) Walker's Mammals of the World. Johns Hopkins University Press: Baltimore.</full-reference>
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        <description>The nocturnal brown hyena roams vast distances on its own each night, around 20 to 35 kilometres, searching for carrion on which to feed (2).  With its exceptional sense of smell the brown hyena can locate carcasses several kilometres away, and without the need to cooperate with any other hyenas, it does not have to share its find unless it brings the food back to the den (6).  Along the Namibian shore, beachcombing for dead fur seals, whales and ocean debris is common (2).  The brown hyena is a poor hunter, but will often make feeble, frequently unsuccessful, attempts to catch any small animal it encounters (3).   It also feeds on invertebrates, eggs and fruit (2), and shrewdly stores any excess food for later use (3).&lt;/p&gt;&lt;p&gt;Despite being solitary when foraging, brown hyenas usually live in extended family groups of four to six individuals, called clans.  A clan defends a territory and all members assist in raising the cubs (3).  Territories are marked by 'pasting' and defecating; behaviours which also form a clever communication system.  Pasting is the act of the hyena depositing secretions from its large anal gland onto grass stalks.  The white blob and thin black smear of paste that is left behind contains information about the hyena &#8211; its identity and the time since the hyena passed by.  This, along with frequently defecating in latrines, ensures that the hyenas of an area have a good idea of what the other clan members are up to (6).&lt;/p&gt;&lt;p&gt;Males in a clan do not usually mate with the clan females, as most of them are related to the females and this would cause inbreeding.  Instead, females usually mate with nomadic males that wander between territories visiting receptive females (3).  The gestation period of the brown hyena is three months, after which up to three blind and deaf young are born.  For the first few months after birthing, the female and her young seclude themselves.  After this period, they rejoin the clan where, in an example of their extraordinary social system, a lactating female may occasionally suckle other cubs than her own, but showing a clear preference toward her own cubs, and all members of the clan help feed the cubs by bringing food back to the den (2) (4) (6).</description>
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      <canonical-form>Animalia</canonical-form>
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      <id type="integer">694</id>
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      <iucn-conservation-status>NOT EVALUATED</iucn-conservation-status>
      <scientific-name>Hyaenidae</scientific-name>
    </taxon-page>
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      <id type="integer">39396</id>
      <canonical-form>Hyaena</canonical-form>
      <iucn-conservation-status>NOT EVALUATED</iucn-conservation-status>
      <scientific-name>Hyaena</scientific-name>
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      <canonical-form>Hyaena brunnea</canonical-form>
      <iucn-conservation-status>Near Threatened (NT)</iucn-conservation-status>
      <scientific-name>&lt;i&gt;Hyaena brunnea&lt;/i&gt; Thunberg, 1820</scientific-name>
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MS Education - Johns Hopkins University
ME Engineering Physics - University of Virginia


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D.Sc., Queen's University, Belfast (1990)
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Visiting Scientist, Chemicals Affecting Insect Behavior Lab, US Agricultural Research Service, 2005-2006
Hammock, J., Vinyard, B., Dickens, J. 2007. Response to host plant odors and aggregation pheromone by larvae of the Colorado potato beetle on a servosphere.  Arthropod-Plant Interactions, 1(1):27-35
PhD, Biological Oceanography, Massachussetts Institute of Technology/Woods Hole Oceangraphic Institution, 2005</credentials>
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Assistant Professor
Baker University
Dept. of Biology
PO Box 65
Baldwin city, Kansas 
66006-0065



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University of Miami
Coral Gables, Fl 33124

AAAS, ICRS, Sigma Xi

http://www.bio.miami.edu/Fac/Sealey.html</credentials>
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Yahnke CJ, 2006. Habitat use and natural history of small mammals in the Paraguayan chaco. Mastozoologia Neotropical, 13(1);103-116.

Yahnke CJ, 2006. Testing optimal foraging theory using bird predation on goldenrod galls. American Biology Teacher, 68(8):471-475.

Yahnke CJ, PL Meserve, TG Ksiazek, and JN Mills. 2001. Patterns of infection with Laguna Negra virus in wild populations of Calomys laucha in the central Paraguayan chaco.  American Journal of Tropical Medicine and Hygiene, 65(6):768-776.

Yahnke CJ, I Gamarra de Fox, and F Colman. 1998. Mammalian species richness in Paraguay: the effectiveness of national parks in preserving biodiversity.  Biological Conservation 84:263-268.

Yahnke CJ, J Unger, BA Lohr, DA Meritt, and W Heuschele. 1997.  Age specific fecundity, litter size, and sex ratio in the chacoan peccary (Catagonus wagneri).  Zoo Biology 16:301-307.

Yahnke, CJ, WE Johnson, E Geffen, D Smith, F Hertel, MS Roy, CF Bonacic, TK Fuller, B Van Valkenburgh, and RK Wayne. 1996. Darwin&#8217;s fox: a distinct endangered species in a vanishing habitat.  Conservation Biology 10:366-375.

Yahnke CJ. 1995.  Metachromism and the insight of Wilfred Osgood: evidence of common ancestry for Darwin&#8217;s fox and the Sechura fox.  Revista Chilena de Historia Natural 68:459-467

American Society of Mammalogists
National Association of Biology Teachers
Sigma XI
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Holdich, Catherine, David M., Noel, Pierre Y., Reynolds, Julian D. and Haffner, Patrick (eds) (2006). Atlas of crayfish in Europe. Museum national d'Histoire naturelle, Paris, 187 pages. Patrimoines naturels, 64.

Hudson, Anne V. and Reynolds, Julian D. (1984). Distribution of Irish intertidal Talitridae. Bulletin of the Irish biogeographical Society, 8, 63-76.

Reynolds, Julian D. (1976). Occurrence of the fresh-water Bryozoan, Cristatella mucedo Cuvier, in British Columbia. Syesis, 9, 365-366.

Smyth, Thomas and Reynolds, Julian D.  (1995). Survival ability of statoblasts of freshwater Bryozoa found in Renvyle Lough, County Galway.  Biology and Environment: Proceedings of the Royal Irish Academy 95B (1), 65-68.

Wickenberg, Maria and Reynolds, Julian D. (2002). A recent Irish record of the woodlouse  Acaeroplastes melanurus (Budde-Lund, 1885) (Isopoda: Porcellionidae), considered to be extinct in the British Isles.  Bulletin of the Irish Biogeographical Society, 26, 60-63.</credentials>
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Author Sponges of the British Isles - A colour guide and working document, 1992 Edition. Ackers, R.G., Moss, D. &amp; Picton, B. E. 1992. Marine Conservation Society, UK. Revised and extended, 2007, Bernard Picton, Christine Morrow &amp; Rob van Soest. PDf and website</credentials>
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