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BiologyAdults occur among rubble or consolidated reef rock of exposed intertidal reef flats and sandy beaches, subject to mild surge. Also near freshwater run-offs (Ref. 48636). They occur in small groups. Feed mainly on benthic algae. Oviparous, distinct pairing during breeding (Ref. 205). Eggs are demersal and adhere to the substrate (Ref. 205). Males guard and aerate the eggs (Ref. 205).