Comprehensive Description

provided by Memoirs of the American Entomological Society
Laccophilus fasciatus rufus Melsheimer, restored name and new status Laccophilus rufus Melsheimer, 1844, p. 28. Type: Harvard University, Museum of Comparative Zoology, number 31234, male, Pennsylvania.
Laccophilus fasciatus, Crotch, 1873, p. 400; Sharp, 1882a, p. 290; Blatchley, 1910. p. 210; Leng, 1920, p. 77; Zimmermann, 1920, p. 18; Zimmerman and Severin, 1957, pp. 29-32; Zimmerman, 1960, pp. 142-3.
DIAGNOSIS. — L. fasciatus can easily be separated from all other irrorated species in the eastern United States by the fascia or blotch on the posterior half of the elytra.
NOMENCLATURAL NOTES. — Say (1823) recognized rufus, but referred to it as a variety of maculosus without giving it a name. This race has long been known as fasciatus; but, as discussed above, the remaining type is the Mexican subspecies and not an individual from the eastern United States. Melsheimer's rufus is restored as valid, since there is no doubt as to the identity of his type in the Museum of Comparative Zoology.
VARIATION. — The principal variation in elytral pattern is shown in Figures 285 and 286. Small specimens with weakened fascias resemble the sympatric proximus. This is individual variation, however; and, apparently, has no geographical significance.
Males and females are very nearly the same size, both averaging about 4.74 mm; and measurements have been combined for each locality.
Apparently, there is considerable regional variation in size of rufus, while pattern and the PW/EL ratio remain fairly uniform throughout the range.
The average values for all four measurements are much the same for three large samples from Indiana, Illinois, and Kansas (Table 2), but three other samples from the eastern United States give a different picture. Sixty-nine specimens from several southeastern states had a mean length of only 4.68 mm, as compared to 4.73 mm for the three midwestern samples and 4.90 mm and 4.81 mm for New York and
Pennsylvania. L. /. rufus appears to be smaller in the southeast than in any other part of the range. The standard deviations were all larger than other comparable samples, which indicates some currently unrecognized difference within southeastern populations.
A cline for all four measurements shows up if the three samples from New York to Indiana are compared. The differences are all significant (95 per cent level and above). This is one of the few clines that have been discovered in this study even though the populations cover large areas. — L. f. rufus ranges from northern Florida and east Texas to Vermont, southern Michigan, and southeastern South Dakota and from the Atlantic Coast to a line approximating the ninety-seventh meridian in South Dakota, Nebraska, Kansas, Oklahoma, and Texas. It normally does not occur in the peninsula of Florida, or the lowlands bordering the Gulf of Mexico; but it has been taken in the vicinity of Mobile Bay. It is probably the most abundant Laccophilus south of the glaciated region in the eastern United States. It intergrades with terminalis from northcentral Kansas to east Texas.
Young (1954) found rufus most commonly in muddy or silty bottomed temporary pools formed in roadside ditches or intermittent streams. These pools were in well-drained soils. It was a pioneer species of recently formed aquatic habitats in upland Florida. Zimmerman (1960) found it in open, unshaded ponds and roadside ditches in clay soils in the Midwest. UNITED STATES OF AMERICA. — ALABAMA. Mobile County. Mobile, 8, J. D. Sherman, Jr. (USNM). Montgomery County. Barachias, 2, vi. 10.24, E. G. Holt (USNM). Russell County. Seale, 1 9, vi.18.54, R. L. Fisher, D. L. Haynes (MCHS). ARKANSAS. Grant County. Poyem, 10 <5 , x.25.48, O. Bryant (CAS). Hempstead County. Hope, 1, ix.7.31, Knoble (BERK); 4 2, iv.3.22, L. Knoble (CAS); 1 c$, vi.11.54, J. W. Green (CAS). CONNECTICUT. Fairfield County. County, 1 2 , vii.— .— , C. H. Roberts (AMNH). DISTRICT OF COLUMBIA. Piney Br., 1, iv.25.05 (USNM); 1, viii.26.06, D. H. demons (USNM). Rock Creek, 6, viii.31.06, D. H. demons (USNM). Washington, 1, vii.17.— ; 1, viii.28.— , Hubbard & Schwarz (USNM); 3, ix.723.06, W. L. McAtee (USNM); 1, vi.6.04, F. Knab (USNM); 1, vii.31.— , H. S. Barker (USNM); 3, iv.28.18, L. L. Buchanan (USNM). District 4, — . — .— (USNM). FLORIDA. Duval County. Jacksonville, 1 $, Van Dyke (CAS). GEORGIA. Chatham County. Savannah, 1, v.17.27, E. W. Mank (CNL). Cobb County. Marietta, 2, iii.5.11 (CNL); 1, iii.5.11 (USNM). Decatur County. Bainbridge, 13,12, vii.15-27.09, J. C. Bradley (CAS); 6, vii.15-27.09; 1, iii.7.10, J. C. Bradley (CNL); County, 2, x.3.38, F. N. Young (CNL). Jefferson County. Wrens, 2, iii.8.11 (CNL); 1, hi. 8. 11 (USNM). Rabun County. Clayton, 1 $, vii. — .10, W. T. Davis (CAS); County, 4 8, 3 2, vii.—.—, C. V. Leng (CAS). ILLINOIS. Alexander County. Olive Branch, 8 3,119, x.8.09, Gerhard (FM). Bond County. Mulberry Grove,
6 3,3 2, x.20.56, JRZ (NMSU). Champaign County. Urbana, 1, vii. 19.07 (USNM); 1 3, iii.— .— (CNG) ; Champaign, 2 3,5 2, xi.5.27, A. T. McClay (DAV). Cook County. (Morton Gr.) Edge Brook, 2, vii. 15. 11, J. D. Sherman, Jr. (USNM). Glencoe, 2, ix.5.04, J. D. Sherman, Jr. (USNM). Palos Pk., 1 2, x. 14.34, W. J. Gerhard (FM). Franklin County. Zeigler, 1, iii. 13.29, J. Karlovich (USNM). Kankakee County. Momence, 1, vii. — . — , Wickham (USNM). Jackson County. Murphysboro, 3 3,12, viii.9.32, W. J. Gerhard (FM). Pike County. Pittsfield, 3 3, viii. 17.46, 13, 22, viii.7.47; 1 3, viii.22.47; 1 2, vii.28.48, B. Cadwell (DAV). Randolph County. Bremen, 1, vii. 19.56, P. J. Spangler, Jr. (USNM). Vermilion County. Oakwood, 1 3, v.5.48, H. T. McClay (DAV). INDIANA. Brown County. Brown County St. Pk., 8 3,7 2, iv.21.56, JRZ (NMSU). Clinton County. Kirklin, 3 m. W., 13,12, ix.12.58; 1 3, 1 2, ix.26.58; 6 3,6 2, x.10.58; 7 3,3 2, x.17.58, JRZ (NMSU). Dubois County. Purdue Exp. Farm, Near Huntingburg, 5 3, 5 2, x.29.55, J. E. Hanegan & JRZ (NMSU). Hamilton County. Westfield,
7 m. N., 3 2, v.4.56, JRZ (NMSU). Huntington County. Huntington, 2 m. N., 7 3, 9 2, x.26.56, JRZ (NMSU). Knox County. Hazelton, 10 m. W., 2 3,22, ix.9.56, JRZ (NMSU) . Lake County. Clarke, 2 2 , viii.7.07, (FM) . Madison County. Alexandria, 5 m. E., 4 3, 3 2, vii. 10.57, JRZ (NMSU). Anderson,2 2,vi.6.57;2 3 , vi.21.57, JRZ (NMSU). Monroe County. Bloomington, 12 3, 12 2, ix.15.55; 6 3, 10 2, ix.20.55; 19 3, 4 2, iii. 13. 56; 22 3, 29 2, iv.26.56; 13,6 2, v.11.56; 13 3, 15 2, vii. 1.56, JRZ (NMSU); 4 m. N., Slough, 12 3, 8 2, vi.18.57, JRZ (NMSU); 4 3,7 2, xii.31.49, F. N. Young (AMNH); 4, xii.31.49, F. N. Young (CNL); ii.9.52, F. N. Young (CNL); 13,1 2, xii.31.49, F. N. Young (FM); 1 3,1 2, same data (ANSP). Morgan County. Waverly, 5 2, vi. 17.57; Morgan-Monroe St. For., 1 3, vii.15.57, JRZ (NMSU). Noble County. Wolfiake, 5 m. S., 13, x.26.56, JRZ (NMSU). Owen County. McCormick's Ck. St. Pk., 3 3,2 2, vii.12.55; Richland Ck., Near Whitehall, 2 3, 2 2, vii.26.55, JRZ (NMSU). Posey County. Mt. Vernon, 5 m. E., 4 3, 7 2, ix.9.56, JRZ (NMSU). Wabash County. Wabash, 3 m. N., 3 3, x.26.56, JRZ (NMSU). IOWA. Decatur County. Leon, 1, vii. 3. 32, F. Andre (TAM). Lee County. Fort Madison, 1 2 (AMNH). Polk County. Herrold, 1, vii. 24.19, E. D. Quirsfeld (CNL). Story County. Ames, 1, v. 11. 32, Floyd Andre (USNM). KANSAS. Atchinson County. Atchinson, 1 3, ix.22.56, J. W. McReynolds (CAS). Cherokee County. Galena, 19 (CNL); 19 (USNM); 33, viii.26.04 (USNM); 6 3,112, Ebb. Crumb (AMNH). Douglas County. Baldwin (City?), 3 3, 4 2, J. C. Bridwell (ORES). Lawrence, 1, x.9.49. J. G. Rozen (BERK); 23 3 , 25 2,
ix.26.— ; x.2.51, P. J. Spangler, Jr. (USNM). Lone Star L., 15 $, 13 2, vi.6-10.52, P. J. Spangler (USNM). County, 2, x.9.48, J. G. Rozen (BERK); 20, iii.20.52; 3, iii.19.52; 1, iii.11.52, P. J. Spangler, Jr. (USNM); 2, F. H. Snow (CNL). Leavenworth County. Lawrence, 7 m. NE, 2, iii.22.52, P. J. Spangler, Jr. (USNM). Ottawa County. County, 1 5, iv.31.33, C. W. Sabrosky (MCHS) . Pottawatomie County. Onaga, 2 (USNM) ; Onega (Onaga?), 1 5, 2 2, F. Crevecoeur (AMNH). Riley County. 1, iii.13.— ; 1, viii.l.— ; 11, x.21.— , Popenoe (USNM). Shawnee County. Topeka, 4 (USNM); 3, Hubbard & Schwarz (USNM); 2 2, Popenoe (MCZ). LOUISIANA. Natchitoches County. Coverdale (AMNH); 2 (USNM). Vowell's Mill, 7 5,92 (CAS). Vernon County. Leesville, 1 2, viii.18.47, H. S. Fitch (DAV). MARYLAND. Baltimore County. 1 5, vii.13.— , F. E. Blaisdell (CAS). Harford County. Edgewood, 2, ix.17.18, H. Dietrich (CNL). Montgomery County. Kensington, 36, vi.2-17.61, J. W. Fitzgerald (USNM). Piney Point, 3, viii.26.46, R. I. Sailer (USNM). Plummers Island, 3, v.6.05; 1, iii.24.07; 3, iv.27.13, W. L. McAtee (USNM). MICHIGAN. Van Buren County. South Haven, 1, viii.l 1.04 (USNM). Wayne County. Detroit, 2, vi.28.— , 2, Hubbard & Schwarz (USNM). MISSISSIPPI. George County. Lucedale, 5, i. 17.32; 4, iii. 27.32, H. Dietrich (CNL). Perry County. Beaumont, 1, iv.19.32, H. Dietrich (CNL). Wilkinson County. Woodville, 1 5, vii.26.21 (CAS). MISSOURI. Boone County. Ashland, 1, iii.29.56, P. J. Spangler, Jr. (USNM). Columbia, 10, iii.21-29.36; 3, v.2.36, W. M. Gordon (CNL); 3, vi.26.— , Wickham (USNM). Columbia, 5 m. S., 1, viii.20.53, M. C. Grabeau (USNM) . Deer Park, 6 m. E., 1, viii.28.53, M. C. Grabeau (USNM) . Cooper County. Booneville, 1 m. E., 6, ix.25.54, P. J. Spangler, Jr. (USNM). Girardeau County. Cape Girardeau, 1, iv.1.56, D. Stout (USNM). Greene County. Springfield, 1, xii. 11.49, E. T. Oxtman (BERK). Mississippi County. Charleston, 1, iv.23.56, P. J. Spangler, Jr. (USNM). St. Louis County. Howard Bend, 7, vi.28. 37; 7, vii.17.37; 2, vii. 22.40; 1, ix.18.40, W. M. Gordon (CNL). Manchester, 1, ix. 15.57, P. J. Spangler, Jr. (USNM). St. Louis, 1, M. Schuster (USNM). Webster Grove, 1, vi.27.33, R. C. Lange, (USNM). Vernon County. Near Nevada, 15,3 2, v.14.58, J. W. McReynolds (CAS). Nevada, 15,12, vi. 11.58, J. W. McReynolds (CAS). County, 1 2, v.30.58, J. W. McReynolds (CAS). NEBRASKA. Lancaster County. Lincoln, 2 5,3 2, x.15.60, W. F. Rapp, Jr. (NMSU). NEW JERSEY. Bergen County. Fort Lee, 7, vii. 30.04, J. D. Sherman, Jr. (USNM). Palisades (Pk.?), 1, iv.19.03, T. D. O'Connor (CNL). County, 4, v. 11.02, Shoemaker (USNM). Camden County. Delair, 3 5,4 2, vii. 16. — , E. L. Dickerson (AMNH). Middlesex County. Avenel, 1 9, v.22.31, Siepmann (FM). Spotswood, 1 2 , C. H. Roberts (AMNH). Morris County. Boonton, 1, vii. 8.01, G. M. Greene (USNM). Ocean County. Ft. Dix, 1 5, vi. 10.43, C. D. Michener (AMNH). Lakehurst, 1, vi.26. 54, D. M. Anderson (CNL); 1, ix.1.01, J. D. Sherman, Jr. (USNM); 4 5, 3 2, vii. 17.23, L. B. Woodruff (AMNH); 1 2, v. 24.— ; 1 5, vii. 3 1 .— , C. H. Roberts ( AMNH) . NEW YORK. New York Vicinity. Staten Is., 1 5, Leng, Van Dyke (CAS); 1; 1, vii.— .91; 17, ix. 15.00, J. D. Sherman, Jr. (USNM); 1 2, ix.27.38; 1 2, x.1.38; 1 2, ix.4.39; 1 2, v.14.39; 15,12, iv.23.39 (FM);2 9, iv.19.— , C. H. Roberts (AMNH); 1 9, ix.27.38 (DAV). N.Y.C., 1 6, 1 9, C. H. Roberts; 1 6, x.24.01 (AMNH). Westchester County. Peekskill, 1, J. D. Sherman, Jr. (USNM). NORTH CAROLINA. Buncombe County. Ashville, L. Ashnoca, 1, iv.29.44, W. E. Hoffman (USNM) ; 1, v.— .44, W. E. Hoffman (USNM). Ashville, 1 9, ix.— .95 (AMNH). Black Mts.,
2 6, v.17.— (AMNH). Madison County. Hot Springs, 1 9 (MCZ). Polk County. Columbus, 1, vi.19.52; 1, vii.14-21.52 (USNM). Guilford County. Greensboro, 1, viii.9. — , F. C. Pratt (USNM). OHIO. Athens County. Athens, 2, x.9-15.49; 3, vi.22-30.50. Waterloo Tnsp., 1, iv.23.50; 2, vi.27.50; 2, vii.6.50, P. J. Spangler, Jr. (USNM). Hamilton County. Cincinnati, 1, viii.17. — , H. Soltau (USNM). Richland County. Mansfield, 1 9 (ORES). Ross County. Londonderry, 2, viii.5.50, P. J. Spangler, Jr. (USNM). Scioto County. Near Friendship, Roosevelt Lake, 3, vii.29.50, P. J. Spangler, Jr. (USNM). Vinton County. Lake Hope, 2 5,1 9, vii.22.52, W. C. Stehr (CAS); 1, x.7.58, W. C. Stehr (NL). Warren County. Mason, 2 m. S., 12 5, 7 9, vi.24.57, J. R. Zimmerman (NMSU). OKLAHOMA. AtokaCounty. Atoka, 1 9 , vi. 13-15.— , Wickham & Fenyes (CAS); 1, vi. 13-15.— , Wickham (USNM). Bryan County. Lake Texoma State Park, 2 5,2 9, vi.24.62, E. Van Tassel (CNA). Marshall County. 1, vi.5.53; 1, viii.5.53 (USNM). Ottawa County. 1 5, vi.5.31, Costner & Davis (MCHS). Payne County. Stillwater, 1 <? , 19, ii. 10.38, K. C. Emerson (MCHS). Pittsburg County. McAlester, 1 $, vi.ll. — , Wickham & Fenyes (CAS); 3, vi.ll.—, Wickham (USNM). PENNSYLVANIA. Adams County . Arendtsville, 1, viii.4.51, P. J. Spangler, Jr. (USNM). Starrs Mill, 1, vi. 15.49, P. J. Spangler, Jr. (USNM). Allegheny County. Pittsburgh,
3 5,49:16, vi.14.— ; 2 9, vii.— .— ; 1 9, viii.— .— ; 1 9, ix.2.— ; 1 9, ix.4.— (CNG). County, 4, iv.5.90, E. A. Klages (CNL); 3 6,89 (CNG). Delaware County. 1 5, v. 18. — (ANSP). Franklin County. Chambersburg, 1, x.l.— ; 2, J. D. Sherman, Jr. (USNM). Juniata County (?). Path (Valley?), 8, J. D. Sherman, Jr. (USNM). Lebanon County. Lebanon, 10, J. D. Sherman. Jr. (USNM). Philadelphia County. Philadelphia, 2 (USNM). Philadelphia? Neck, 15,2 9, v. 12.—, Blaisdell (CAS). Phila. Neck, 2 5, Laurent (AMNH); 2 5,7 9, v.4.— ; 1 6,4 9,ix.l3.— (ANSP): 1: 1, vii. 15.04; 1, v.2.— , Kaeber (USNM): 2, vi.9.00; 1, ix.13.— , G. M. Greene; 2, P. Lauren (USNM). Westmoreland County. Jeannette, 1 6,5 9, vii. — . — ; 1 5, vii. 3. — : 1 9, vii.4.— ; 6 9, vii. 14.— ; 1 9, vi.21.— ; 16, 19, vi.24.— ; 2 5, 3 9, vi.27.— (CNG). SOUTH CAROLINA. Beaufort County. Beaufort, 4 5, 3 9; 2 5, iv.12.92; 1 5, iv.18.92, G. D. Bradford (AMNH). Kershaw County. Camden, 4 5, 1 9, vi.23.29, P. J. Darlington, Jr. (MCZ). Pickens County. Clemson, 2, xi. 25.27, D. Dunavan (USNM) . TENNESSEE. Davidson County. Nashville, 1, viii.4-15.97, Wickham (USNM): 1, Osburn (USNM). Fentress County. Ailardt, 1 5 , viii. 17.22, T. H. Hubbell (MCHSU). McMinn County. Athens, 1 9, vi.24.63, A. R. Moldenke & J. A. Woods (AMNH). Shelby County. Memphis, 1 6, v. 16.1 2, Blaisdell (CAS). TEXAS. Brazos County. College Station, 1, iii.31.35, H. J. Reinhard; 1, vii.19.59, H. R. Burke (TAM). Collin County. Piano, 1, vii.— .07, E. S. Tucker (USNM). Harris County.
Houston, 5, vi.15.32, Harwood (CNL). Harrison County. Karnack, 4 5,7 9, vi.17.58, R. L. Fischer, J. Eichmeier & R. A. Schneibner (MCHS). Panola County. Carthage, 1 $, v.8.52, M. Cazier, W. Gertsch & R. Schrammel (AMNH). VIRGINIA. Albemarle County. Ivy (Depot?), 5 9, vi.— .33, F. E. Winters (CAS). Alexander County. 1, vi.18.10; 1, vii.7.08 (USNM). Bath County. Warm Springs, 1 9, x.l. — , Leng & Van Dyke (CAS). Fairfax County. Great Falls, 2, i.9.06; 1, iii.17.07, D. H. demons (USNM). County, 2, ix.20-22.27, Shoemaker (USNM). Henrico County. Richmond, 4 $, Slossom (AMNH); 1 9 (MCZ). Nansemond County. Suffolk, 10 $, 6 9, vi.27.1876, Dimwock (MCZ). Nelson County. 5, viii.6.16, W. Robinson (USNM). Nottoway County. Camp Pickett, 2 3,1 9 , v. 1.52, B. Adelson (CAS) . Spotsylvania County. Fredericksburg, 1, iii.16.89; 3, iii.15.91; 2, x.4.97; 3, iii. 19-26.99, W. D. Richardson (USNM); 3, ix.— .98, J. D. Sherman, Jr. (USNM); 1 $, 1 9, ii. — .22 (AMNH). VERMONT. Bennington County. County, 15,19, vii. — .94, C. H. Roberts (AMNH); 1 9 (ASNP). WEST VIRGINIA. Greenbriar County. White Sulphur Springs, 1 9, vii. — . — , A. Fenyes (CAS); 10, viii.23.10, W. Robinson (USNM). Jackson County. Ripley, 3, vi.25.30, F. N. Musgrave (CNL). Jefferson County. Harper's Ferry, 4, ix.12.17, J. E. Benedict, Jr. (USNM). Kanawha County. County, 2(5,3$, vii.— .35 (CNG).
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bibliographic citation
Zimmerman, J.R. 1970. A Taxonomic Revision of the aquatic beetle genus Laccophilus (Dytiscidae) of North America. Memoirs of the American Entomological Society vol. 26. Philadelphia, USA