Amambay, Boquerón , Canindeyú , Central, Concepción , Cordillera, Itapúa , Misiones, Ñeembucú , Paraguarí , Pte. Hayes (ALWC, IFML, INBP, LACM, MHNG, NHMW). Literature records: Cordillera(Forel 1906, Forel 1909).