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Comprehensive Description

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Notiphila (Dichaeta) uliginosa Haliday

Notiphila uliginosa Haliday, 1839:222.

Notiphila (Agrolimna) olivacea Cresson [in part], 1917:52.

TYPE-MATERIAL.—I have not located Haliday's syntype series of this species, but Haliday's original description does list the following localities for the syntypes (all in Ireland or Northern Ireland): (1) marsh between Glengariff and Adrighoule (County Cork); (2) banks of the Shannon at Tarbert (County Kerry, Shannon River); (3) shores of Belfast Lough (Northern Ireland). All of the localities for the syntype series are brackish-water marshes along bays or rivers leading to the ocean. I have examined European specimens from Denmark and Sweden.

DIAGNOSIS.—N. uliginosa is a variable species, whose specimens closely resemble other members of the scalaris group, especially N. quadrisetosa. Specimens may be distinguished from all related species by characters of the male postabdomen, especially the shape of the hypandrial process (Figures 197–203). Externally, specimens of N. uliginosa differ from those of N. olivacea in the extent and degree of coloration contrast between the darkened abdominal fascia and the olivaceous to gray background color. Usually the color of the fascia of N. uliginosa members do not contrast distinctly but merge gradually together, and generally the fascia are also more extensive. The melanic form of N. uliginosa members may be differentiated from those of N. quadrisetosa by the absence of a median, bifurcating, mesonotal stripe. Often the facial color is grayer in the melanic form of N. uliginosa, but there is considerable overlap in this character with specimens of N. quadrisetosa.

DESCRIPTION.—Medium-sized to moderately large shore flies, length 3.46 to 4.53 mm; with yellowish brown to olivaceous background coloration and some brown markings.

Head: Head ratio 1:0.73; postfrons ratio 1:0.61; frons mostly dark brown with some darker, charcoal coloration; median triangular area and lateral margins concolorous, lighter in color, tending to be more yellowish. Paravertical bristle large, approximately one-half to one-third longer than largest postocellar setae; 2 pair of proclinate, fronto-orbital setae. Antennal segments entirely dark, dark brown to black; arista with 7–9 dorsal branches. Face dusted yellow; prefrons ratio 1:0.74; facial setae hairlike, numbering 6–7 and usually extending dorsally past midheight of face. Eye ratio 1:0.78; eye-to-cheek ratio 1:0.21. Gena fairly narrow, concolorous with face anteriorly, becoming grayer posteriorly; genal bristle distinct, subequal to paravertical bristle. Maxillary palpus dark, brownish black.

Thorax: Mesonotum slightly darker in coloration than pleural areas but not distinctly contrasting, immaculate. Scutellum approximately concolorous with mesonotum. Mesopleuron generally immaculate; remaining pleural areas tending to become lighter ventrally but not distinctly grayed or contrasting with dorsal portions. Femora dark, dusted gray; tibiae black, also with some gray dusted areas; front tarsomeres mostly blackish but with extensive pale areas at apices and beneath, middle and hind tarsi pale. Setal fascicle of hind basitarsus pale, yellowish orange. Wing ratio 1:0.42; costal vein ratio 1:0.50; M1+2 vein ratio 1:1.1.

Abdomen: Abdomen ratio of males 1:0.64; length of fourth tergum to fifth tergum ratio of males 1:0.97; fifth tergum ratio of males 1:0.61. Coloration of terga mostly olivaceous, darker areas on third, fourth, and fifth terga rather diffuse, not distinctly contrasting with background color, gradually merging. Male terminalia as in Figures 194–203.

Melanic form. As above except as follows. Face often gray but usually dusted yellow. Legs entirely dark, usually black except for tarsal brushes and setal fascicle of hind basitarsus which are pale. Abdominal fascia brown, distinct on terga 3–5; fifth tergum almost entirely dark. The general coloration also tends to be more brown rather than olivaceous.

SPECIMENS EXAMINED (518).—CANADA: ALBERTA: Banff (CNC); 20.9 km N Banff-Jasper Highway (CNC); Cooking Lake (ANSP); Carmangay-Little Bow River (USNM); Edmonton (CAS); Full Lake (ANSP); Gull Lake (CNC); McMurray (CNC); Nordegg (ANSP, CNC); Waterton Lakes (CNC). BRITISH COLUMBIA: Atlin (CNC); Clinton (CNC); King Salmon Lake (CNC); Lakelse Hot Spring (CNC); Lakelse near Terrace (CNC); Vernon (CNC). MANITOBA: Brandon (CNC); 8.1 km SW Shilo (CNC); Wabawden (CNC). NORTHWEST TERRITORIES: Fort Smith (CNC); Norman Wells (CNC). ONTARIO: Marmora (CNC). QUEBEC: Brome Lake (CNC); Covey Hill (CNC); Lac Mondor-Ste. Flore (CNC); La Trappe (AMNH, FSCA, MSU); Missisquoi Bay (CNC); Rupert House (CNC); Saint Chrysostome (CNC); Saint Placide (AMNH). SASKACHEWAN: Assiniboia (CNC); Attons Lake-Cut Knife (CNC); Cut Knife (CNC); Lisieux (CNC); Rutland Station (CNC); Val Marie (CNC); Willows (CNC). YUKON TERRITORY: Watson Lake (CNC). UNITED STATES: ALASKA: Anchorage (ANSP); Auke Bay (WSU); Unalakleet (CNC); Valdez, tidal flats (USNM). CALIFORNIA: Lassen Co., 66 km NW Susanville (WNM); Marin Co., 24.2 km NW Olema (UCB); Modoc Co., 6.4 km W Alturas (WMN); Lookout (WMN), Surprise Valley-Fort Bidwell (UCB); Nevada Co., 1.6 km NW Hobart Mills (WNM), 4 km NE Hobart Mills (WNM). COLORADO: Boulder Co., Nederland (ANSP, CSU, KU, USNM, WSU); Clear Creek Co., Echo Lake-Mount Evans (CNC); Park Co., Kenosha Pass (CSU); Saguache Co., Saguache (CSU). ILLINOIS: Jo Daviess Co., Galena (INHS): Lake Co., Lake Forest (CU). MICHIGAN: Mackinac Co. (MSU); Manistee Co., Manistee (ANSP). MASSACHUSETTS: Cook Co., Grand Marais (UMN), Marais (UMN); Marshall Co. (UMN). MONTANA: Flathead Co., 1.6 km W Bigfork (KSU, USNM), 12.9 km S West Glacier (KSU); Glacier Co., 3.2 km E Babb (KSU, USNM), NW Browning (KSU, USNM); Lake Co., 3.7 km E.) Bigfork (WNM), Dayton (ANSP), 32.2 km S Swan Lake (KSU). NEW MEXICO: Sandoval Co., Jemez Springs (ANSP). OREGON: Benton Co., 19.3 km S Corvallis-McFadden Pond (WNM); Grant Co., 0.8 km S Senaca (WNM); Harney Co., 25.8 km N Burns (WNM), 3.2 km E Burns (WNM), S shore Harney Lake (WNM); Klamath Co., Aspen Lake (WNM); Lake Co., Ana Reservoir (WNM), 6.9 km NW Paisley (WNM), 7.7 km N Summer Lake (WNM), Warner Canyon (WNM); Linn Co., 33.8 km SE Idanha (WNM); Wheeler Co., 29.8 km E Mitchell (WNM). SOUTH DAKOTA: Custer Co., 11.3 km W Custer (WNM), Flynn Creek 12.9 km N Pringle (CNC); Grant Co., Big Stone City (USNM); Meade Co., Piedmont (KU); Stanley Co., Fort Pierre (ANSP). UTAH: Utah Co., Provo Canyon (UMN). VERMONT: Windham Co., Jacksonville-Laurel Lake (UMN). WASHINGTON: Grant Co., Grand Coulee-Columbia River (WSU); Ferry Co., 24.2 km W Kettle Falls (UN, USNM); Spokane Co., Liberty Lake (USNM); Whitman Co., Johnson (WSU)

GEOGRAPHIC DISTRIBUTION (Figure 204).—Notiphila uliginosa is distributed primarily in the West, where it is found as far north as Alaska, Yukon Territory, and the Northwest Territories and eastward as far as Vermont. Collection dates are from 15 May to 28 August.
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bibliographic citation
Mathis, Wayne Neilsen. 1979. "Studies of Notiphilinae (Diptera: Ephydridae), I: Revision of the Neartic Species of Notiphila Fallen, Excluding the Caudata Group." Smithsonian Contributions to Zoology. 1-111. https://doi.org/10.5479/si.00810282.287