Image of Exocelina alexanderi Shaverdo, Hendrich & Balke

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Exocelina

Scientific Names

Preferred Names

Genus recognized by EOL Dynamic Hierarchy April 2022

Reference taxon from Catalogue of Life in Species 2000 & ITIS Catalogue of Life

Genus recognized by wikipedia русскую Википедию, wikipedia SV, EOL Dynamic Hierarchy 1.1, EOL Dynamic Hierarchy 1.1, Australia Species List, New Caledonia Species List, Indian Ocean Species List, NCBI, wikipedia MIN, wikidata_hierarchy tar gz, wikipedia ES, wikipedia EN, Wikidata Common Names, Wikimedia Commons, Barcode of Life Data Systems, and Dynamic Hierarchy Version 2.2 - Test

Genus recognized by GBIF classification, Catalogue of Life 7 February 2023, and Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy

Alternative Names

Heterotypic synonym according to GBIF classification

Heterotypic synonym according to GBIF classification

Heterotypic synonym according to GBIF classification

Heterotypic synonym according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Doubtful according to GBIF classification

Invalid according to Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy

Harvest ancestor according to GBIF national node type records France, iNaturalist, Simpson et al 2020, and TreatmentBank

Invalid according to Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy

Synonym according to Catalogue of Life 7 February 2023 and Catalogue of Life 7 February 2023

Common Names

There are no common names associated with this taxon.

Life

Cellular Organisms

Eukaryota

Opisthokonta

Metazoa

Bilateria

Protostomia

Ecdysozoa

Arthropoda

Pancrustacea

Hexapoda

Insecta

Pterygota

Neoptera

Endopterygota

Coleoptera

Adephaga

Dytiscidae

Exocelina (this page)

Exocelina abdita (Balke, Watts, Cooper, Humphreys & Vogler 2004)

Exocelina aipo (Balke 1998)

Exocelina aipomek (Balke 1998)

Exocelina allerbergeri Wewalka, Balke & Hendrich 2010

Exocelina ascendens (Balke 1998)

Exocelina astrophallus (Balke 1998)

Exocelina atowaso (Shaverdo, Sagata & Balke 2005)

Exocelina atra (Sharp 1882)

Exocelina atrata (J. Balfour-Browne 1939)

Exocelina atripennis (J. Balfour-Browne 1939)

Exocelina aubei (Montrouzier 1860)

Exocelina australiae (Clark 1863)

Exocelina australis (Clark 1863)

Exocelina bacchusi (Balke 1998)

Exocelina bagus (Balke & Hendrich ex Balke 2001)

Exocelina barbarae Wewalka, Balke & Hendrich 2010

Exocelina bimaculata (Perroud & Montrouzier 1864)

Exocelina boulevardi (Watts 1978)

Exocelina broschii (Balke 1998)

Exocelina brownei (Guignot 1942)

Exocelina brunoi Wewalka, Balke & Hendrich 2010

Exocelina burwelli Wewalka, Balke & Hendrich 2010

Exocelina casuarina (Balke & Hendrich 1998)

Exocelina charlottae Wewalka, Balke & Hendrich 2010

Exocelina commatifera (Heller 1916)

Exocelina creuxorum Wewalka, Balke & Hendrich 2010

Exocelina damantiensis (Balke 1998)

Exocelina danae (Balke 1998)

Exocelina desii (Balke 1999)

Exocelina elongatula (W. J. MacLeay 1871)

Exocelina erteldi (Balke 1998)

Exocelina ferruginea (Sharp 1882)

Exocelina feryi Wewalka, Balke & Hendrich 2010

Exocelina flammi Wewalka, Balke & Hendrich 2010

Exocelina fume (Balke 1998)

Exocelina gapa (Watts 1978)

Exocelina gaulorum Wewalka, Balke & Hendrich 2010

Exocelina gelima Wewalka, Balke & Hendrich 2010

Exocelina glypta (Guignot 1955)

Exocelina gracilis (Sharp 1882)

Exocelina heidiae (Balke 1998)

Exocelina hintelmannae (Shaverdo, Sagata & Balke 2005)

Exocelina inexspectata Wewalka, Balke & Hendrich 2010

Exocelina interrupta (Perroud & Montrouzier 1864)

Exocelina jaseminae (Balke 1998)

Exocelina jeannae Wewalka, Balke & Hendrich 2010

Exocelina kainantuensis (Balke 2001)

Exocelina karmurensis (Balke 1998)

Exocelina ketembang (Balke 1998)

Exocelina koghis Wewalka, Balke & Hendrich 2010

Exocelina kolleri Wewalka, Balke & Hendrich 2010

Exocelina larsoni (Balke 1998)

Exocelina leae Wewalka, Balke & Hendrich 2010

Exocelina lilianae Wewalka, Balke & Hendrich 2010

Exocelina maculata (Sharp 1882)

Exocelina madangensis (Balke 2001)

Exocelina manfredi (Balke 1998)

Exocelina marinae (Shaverdo, Sagata & Balke 2005)

Exocelina me (Balke 1998)

Exocelina melanaria (Sharp 1882)

Exocelina messeri (Balke 1999)

Exocelina miriae (Balke 1998)

Exocelina monae (Balke 1998)

Exocelina monteithi Wewalka, Balke & Hendrich 2010

Exocelina munaso (Shaverdo, Sagata & Balke 2005)

Exocelina nielsi Wewalka, Balke & Hendrich 2010

Exocelina niklasi Wewalka, Balke & Hendrich 2010

Exocelina nomax (J. Balfour-Browne 1939)

Exocelina novaecaledoniae (J. Balfour-Browne 1939)

Exocelina ouin Wewalka, Balke & Hendrich 2010

Exocelina parvula (Boisduval 1835)

Exocelina patepensis (Balke 1998)

Exocelina perfecta (Sharp 1882)

Exocelina poellabauerae Wewalka, Balke & Hendrich 2010

Exocelina polita (Sharp 1882)

Exocelina punctipennis (Lea 1899)

Exocelina rasilis (Lea 1899)

Exocelina rasjadi Watts & Humphreys 2009

Exocelina remyi Wewalka, Balke & Hendrich 2010

Exocelina rivulus (Balke 1998)

Exocelina rotteri Wewalka, Balke & Hendrich 2010

Exocelina rufa (Balke 1998)

Exocelina sanctimontis (Balke 1998)

Exocelina schoelleri Wewalka, Balke & Hendrich 2010

Exocelina shizong (Balke & Bergsten 2003)

Exocelina simoni Wewalka, Balke & Hendrich 2010

Exocelina simplex (Clark 1863)

Exocelina staneki Wewalka, Balke & Hendrich 2010

Exocelina subjecta (Sharp 1882)

Exocelina takime (Balke 1998)

Exocelina talaki (Balke 1998)

Exocelina tarmluensis (Balke 1998)

Exocelina ullrichi (Balke 1998)

Exocelina vladimiri (Shaverdo, Sagata & Balke 2005)

Acilius

Aethionectes

Africodytes

Africophilus

Agabinus

Agabus

Agaporomorphus

Aglymbus

Allomatus

Allopachria

Amarodytes

Andex

Anginopachria

Anisomeria

Bidessodes

Canthyporus

Carabdytes

Carabhydrus

Coelhydrus

Colymbetes

Coptotomus

Crinodessus

Deronectes

Derovatellus

Dimitshydrus

Ereboporus

Eretes

Fontidessus

Graphoderus

Heroceras

Herophydrus

Heterosternuta

Hoperius

Huxelhydrus

Hydaticus

Hyderodes

Hydrodessus

Hydrodytes

Hydroglyphus

Hydronebrius

Hydroporus

Hydrotrupes

Hygrotus

Hyphoporus

Hyphydrus

Hypodessus

Iberoporus

Ilybiosoma

Ilybius

Japanolaccophilus

Kuschelydrus

Laccodytes

Lacconectus

Laccophilus

Laccornellus

Laccornis

Laccosternus

Leiodytes

Limbodessus

Madaglymbus

Megadytes

Melanodytes

Metaporus

Methles

Microdessus

Microhydrodytes

Napodytes

Nebrioporus

Neobidessus

Neoclypeodytes

Neoscutopterus

Neptosternus

Notaticus

Onychohydrus

Oreodytes

Pachynectes

Paroster

Peschetius

Philaccolilus

Phreatodessus

Platambus

Platydytes

Platynectes

Primospes

Rhithrodytes

Rugosus

Sandracottus

Scarodytes

Senilites

Siamoporus

Siettitia

Spencerhydrus

Sternopriscus

Stictonectes

Tepuidessus

Trichonectes

Trogloguignotus

Typhlodessus

Yola

Yolina

78 additional siblings truncated for brevity.

See the resource file for a full list.

Biota

Animalia

Arthropoda

Insecta

Coleoptera

Dytiscoidea

Dytiscidae Leach 1815

Copelatinae Branden 1885

Copelatini Branden 1885

Exocelina Broun 1886 (this page)

Exocelina abdita (Balke, Watts, Cooper, Humphreys & Vogler 2004)

Exocelina adelbertensis Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina aipo (Balke 1998)

Exocelina aipomek (Balke 1998)

Exocelina akameku Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina alexanderi Shaverdo, Hendrich & Balke 2012

Exocelina allerbergeri Wewalka, Balke & Hendrich 2010

Exocelina amabilis Shaverdo & Balke ex Shaverdo et al. 2020

Exocelina ambua Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina andakombensis Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina anggiensis Shaverdo, Hendrich & Balke 2012

Exocelina apistefti Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina arfakensis Shaverdo, Hendrich & Balke 2012

Exocelina ascendens (Balke 1998)

Exocelina aseki Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina astrophallus (Balke 1998)

Exocelina athesphati Shaverdo, Surbakti, Sumoked & Balke 2020

Exocelina atowaso (Shaverdo, Sagata & Balke 2005)

Exocelina atra (Sharp 1882)

Exocelina atrata (J. Balfour-Browne 1939)

Exocelina atripennis (J. Balfour-Browne 1939)

Exocelina aubei (Montrouzier 1860)

Exocelina australiae (Clark 1863)

Exocelina australis (Clark 1863)

Exocelina bacchus Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina bagus (Balke & Hendrich ex Balke 2001)

Exocelina baliem Shaverdo, Hendrich & Balke 2013

Exocelina barbarae Wewalka, Balke & Hendrich 2010

Exocelina bewani Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina bewaniensis Shaverdo, Menufandu & Balke 2014

Exocelina bifida Shaverdo, Hendrich & Balke 2012

Exocelina bimaculata (Perroud & Montrouzier 1864)

Exocelina bismarckensis Shaverdo & Balke 2014

Exocelina boulevardi (Watts 1978)

Exocelina brahminensis Shaverdo, Hendrich & Balke 2012

Exocelina brazza Shaverdo & Balke ex Shaverdo et al. 2020

Exocelina broschii (Balke 1998)

Exocelina brownei (Guignot 1942)

Exocelina brunoi Wewalka, Balke & Hendrich 2010

Exocelina bundiensis Shaverdo, Hendrich & Balke 2012

Exocelina burwelli Wewalka, Balke & Hendrich 2010

Exocelina casuarina (Balke & Hendrich 1998)

Exocelina charlottae Wewalka, Balke & Hendrich 2010

Exocelina cheesmanae (J. Balfour-Browne 1939)

Exocelina commatifera (Heller 1916)

Exocelina craterensis Shaverdo & Balke 2014

Exocelina creuxorum Wewalka, Balke & Hendrich 2010

Exocelina cyclops Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina damantiensis (Balke 1998)

Exocelina danae (Balke 1998)

Exocelina desii (Balke 1999)

Exocelina edeltraudae Shaverdo, Hendrich & Balke 2012

Exocelina ekari Shaverdo, Hendrich & Balke 2012

Exocelina ekpliktiki Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina elongatula (W. J. MacLeay 1871)

Exocelina eme Shaverdo, Hendrich & Balke 2012

Exocelina erteldi (Balke 1998)

Exocelina evelyncheesmanae Shaverdo, Hendrich & Balke 2012

Exocelina ferruginea (Sharp 1882)

Exocelina feryi Wewalka, Balke & Hendrich 2010

Exocelina flammi Wewalka, Balke & Hendrich 2010

Exocelina foja Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina fume (Balke 1998)

Exocelina gapa (Watts 1978)

Exocelina garana Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina gaulorum Wewalka, Balke & Hendrich 2010

Exocelina gelima Wewalka, Balke & Hendrich 2010

Exocelina glypta (Guignot 1955)

Exocelina gorokaensis Shaverdo & Balke 2014

Exocelina gracilis (Sharp 1882)

Exocelina haia Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina hansferyi Shaverdo, Hendrich & Balke 2012

Exocelina heidiae (Balke 1998)

Exocelina herowana Shaverdo & Balke 2014

Exocelina hintelmannae (Shaverdo, Sagata & Balke 2005)

Exocelina hudsoni Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina ibalimi Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina inexspectata Wewalka, Balke & Hendrich 2010

Exocelina injiensis Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina interrupta (Perroud & Montrouzier 1864)

Exocelina iratoi Shaverdo, Wild, Sumoked & Balke 2017

Exocelina irianensis Shaverdo, Hendrich & Balke 2012

Exocelina jaseminae (Balke 1998)

Exocelina jeannae Wewalka, Balke & Hendrich 2010

Exocelina jimiensis Shaverdo & Balke 2014

Exocelina kabwumensis Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina kailaki Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina kainantuensis (Balke 2001)

Exocelina kakapupu Shaverdo, Hendrich & Balke 2012

Exocelina karmurensis (Balke 1998)

Exocelina keki Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina ketembang (Balke 1998)

Exocelina kinibeli Shaverdo & Balke 2014

Exocelina kisli Shaverdo & Balke 2014

Exocelina knoepfchen Shaverdo, Hendrich & Balke 2012

Exocelina kobau Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina koghis Wewalka, Balke & Hendrich 2010

Exocelina kolleri Wewalka, Balke & Hendrich 2010

Exocelina koroba Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina kowalskii Shaverdo & Balke 2019

Exocelina ksionseki Shaverdo & Balke 2014

Exocelina kumulensis Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina larsoni (Balke 1998)

Exocelina leae Wewalka, Balke & Hendrich 2010

Exocelina lembena Shaverdo & Balke 2014

Exocelina likui Shaverdo, Wild, Sumoked & Balke 2017

Exocelina lilianae Wewalka, Balke & Hendrich 2010

Exocelina maculata (Sharp 1882)

Exocelina manfredi (Balke 1998)

Exocelina manokwariensis Shaverdo, Panjaitan & Balke 2016

Exocelina mantembu Shaverdo & Balke 2014

Exocelina marawaka Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina marinae (Shaverdo, Sagata & Balke 2005)

Exocelina me (Balke 1998)

Exocelina mekilensis Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina melanaria (Sharp 1882)

Exocelina mendiensis Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina menyamya Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina messeri (Balke 1999)

Exocelina mianminensis Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina michaelensis Shaverdo & Balke 2014

Exocelina mimika Shaverdo & Balke ex Shaverdo et al. 2020

Exocelina miriae (Balke 1998)

Exocelina monae (Balke 1998)

Exocelina mondmillensis Shaverdo, Sagata & Balke 2016

Exocelina monteithi Wewalka, Balke & Hendrich 2010

Exocelina morobensis Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina munaso (Shaverdo, Sagata & Balke 2005)

Exocelina nehoue Balke, Hájek, Hendrich & Wewalka 2014

Exocelina nielsi Wewalka, Balke & Hendrich 2010

Exocelina niklasi Wewalka, Balke & Hendrich 2010

Exocelina nomax (J. Balfour-Browne 1939)

Exocelina novaecaledoniae (J. Balfour-Browne 1939)

Exocelina oceai Shaverdo, Hendrich & Balke 2012

Exocelina oiwa Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina okapa Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina okbapensis Shaverdo & Balke ex Shaverdo et al. 2017

Exocelina oksibilensis Shaverdo, Surbakti, Warikar & Balke ex Shaverdo et al. 2019

Exocelina oraia Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina ouin Wewalka, Balke & Hendrich 2010

Exocelina parvula (Boisduval 1835)

Exocelina perfecta (Sharp 1882)

Exocelina pinocchi Shaverdo & Balke 2014

Exocelina piusi Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina poellabauerae Wewalka, Balke & Hendrich 2010

Exocelina polita (Sharp 1882)

Exocelina posmani Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina pseudoastrophallus Shaverdo & Balke 2014

Exocelina pseudobifida Shaverdo & Balke 2014

Exocelina pseudoedeltraudae Shaverdo & Balke 2014

Exocelina pseudoeme Shaverdo & Balke 2014

Exocelina pseudofume Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina pseudojaseminae Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina pseudomarinae Shaverdo, Sagata & Balke 2016

Exocelina pseudopusilla Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina pseudosoppi Shaverdo, Hendrich & Balke 2012

Exocelina pui Shaverdo, Wild, Sumoked & Balke 2017

Exocelina pulchella Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina pulukensis Shaverdo, Wild, Sumoked & Balke 2017

Exocelina punctipennis (Lea 1899)

Exocelina pusilla Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina ransikiensis Shaverdo, Panjaitan & Balke 2016

Exocelina rasilis (Lea 1899)

Exocelina rasjadi Watts & Humphreys 2009

Exocelina remyi Wewalka, Balke & Hendrich 2010

Exocelina riberai Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina rotteri Wewalka, Balke & Hendrich 2010

Exocelina rufa (Balke 1998)

Exocelina saltusholmesensis Watts, Hendrich & Balke 2016

Exocelina saudaunensis Shaverdo & Balke 2014

Exocelina schoelleri Wewalka, Balke & Hendrich 2010

Exocelina shizong (Balke & Bergsten 2003)

Exocelina sima Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina simbaiarea Shaverdo & Balke 2014

Exocelina simbaiensis Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina simbaijimi Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina simoni Wewalka, Balke & Hendrich 2010

Exocelina simplex (Clark 1863)

Exocelina skalei Shaverdo & Balke 2014

Exocelina soppi Shaverdo, Hendrich & Balke 2012

Exocelina staneki Wewalka, Balke & Hendrich 2010

Exocelina subjecta (Sharp 1882)

Exocelina sugayai Balke & Ribera 2020

Exocelina sumokedi Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina tabubilensis Shaverdo & Balke 2014

Exocelina takime (Balke 1998)

Exocelina talaki (Balke 1998)

Exocelina tariensis Shaverdo & Balke 2014

Exocelina tekadu Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina tomhansi Shaverdo, Wild, Sumoked & Balke 2017

Exocelina tsinga Shaverdo, Surbakti, Sumoked & Balke 2020

Exocelina ullrichi (Balke 1998)

Exocelina unipo Shaverdo, Hendrich & Balke 2012

Exocelina utowaensis Shaverdo, Hendrich & Balke 2012

Exocelina varirata Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina vladimiri (Shaverdo, Sagata & Balke 2005)

Exocelina vovai Shaverdo & Balke 2014

Exocelina waaf Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina waigeoensis Shaverdo, Hendrich & Balke 2012

Exocelina wannangensis Shaverdo & Balke 2014

Exocelina warahulenensis Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina warasera Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina wareaga Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina weylandensis Shaverdo, Hendrich & Balke 2012

Exocelina wigodukensis Shaverdo, Wild, Sumoked & Balke 2017

Exocelina woitapensis Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina wondiwoiensis Shaverdo, Hendrich & Balke 2012

Exocelina yoginofi Shaverdo & Balke ex Shaverdo et al. 2018

Agaporomorphus Zimmermann 1921

Aglymbus Sharp 1880

Capelatus Turner & Bilton 2015

Copelatus Erichson 1832

Lacconectus Motschulsky 1855

Liopterus Dejean 1833

Madaglymbus Shaverdo & Balke ex Shaverdo, Monaghan, Lees, Ranaivosolo & Balke 2008

Life

Cellular Organisms

Eukaryota

Opisthokonta

Metazoa

Bilateria

Protostomia

Ecdysozoa

Arthropoda

Pancrustacea

Hexapoda

Insecta

Pterygota

Neoptera

Endopterygota

Coleoptera

Adephaga

Dytiscidae

Exocelina (this page)

Exocelina abdita (Balke, Watts, Cooper, Humphreys & Vogler 2004)

Exocelina aipo (Balke 1998)

Exocelina aipomek (Balke 1998)

Exocelina allerbergeri Wewalka, Balke & Hendrich 2010

Exocelina ascendens (Balke 1998)

Exocelina astrophallus (Balke 1998)

Exocelina atowaso (Shaverdo, Sagata & Balke 2005)

Exocelina atra (Sharp 1882)

Exocelina atrata (J. Balfour-Browne 1939)

Exocelina atripennis (J. Balfour-Browne 1939)

Exocelina aubei (Montrouzier 1860)

Exocelina australiae (Clark 1863)

Exocelina australis (Clark 1863)

Exocelina bacchusi (Balke 1998)

Exocelina bagus (Balke & Hendrich ex Balke 2001)

Exocelina barbarae Wewalka, Balke & Hendrich 2010

Exocelina bimaculata (Perroud & Montrouzier 1864)

Exocelina boulevardi (Watts 1978)

Exocelina broschii (Balke 1998)

Exocelina brownei (Guignot 1942)

Exocelina brunoi Wewalka, Balke & Hendrich 2010

Exocelina burwelli Wewalka, Balke & Hendrich 2010

Exocelina casuarina (Balke & Hendrich 1998)

Exocelina charlottae Wewalka, Balke & Hendrich 2010

Exocelina commatifera (Heller 1916)

Exocelina creuxorum Wewalka, Balke & Hendrich 2010

Exocelina damantiensis (Balke 1998)

Exocelina danae (Balke 1998)

Exocelina desii (Balke 1999)

Exocelina elongatula (W. J. MacLeay 1871)

Exocelina erteldi (Balke 1998)

Exocelina ferruginea (Sharp 1882)

Exocelina feryi Wewalka, Balke & Hendrich 2010

Exocelina flammi Wewalka, Balke & Hendrich 2010

Exocelina fume (Balke 1998)

Exocelina gapa (Watts 1978)

Exocelina gaulorum Wewalka, Balke & Hendrich 2010

Exocelina gelima Wewalka, Balke & Hendrich 2010

Exocelina glypta (Guignot 1955)

Exocelina gracilis (Sharp 1882)

Exocelina heidiae (Balke 1998)

Exocelina hintelmannae (Shaverdo, Sagata & Balke 2005)

Exocelina inexspectata Wewalka, Balke & Hendrich 2010

Exocelina interrupta (Perroud & Montrouzier 1864)

Exocelina jaseminae (Balke 1998)

Exocelina jeannae Wewalka, Balke & Hendrich 2010

Exocelina kainantuensis (Balke 2001)

Exocelina karmurensis (Balke 1998)

Exocelina ketembang (Balke 1998)

Exocelina koghis Wewalka, Balke & Hendrich 2010

Exocelina kolleri Wewalka, Balke & Hendrich 2010

Exocelina larsoni (Balke 1998)

Exocelina leae Wewalka, Balke & Hendrich 2010

Exocelina lilianae Wewalka, Balke & Hendrich 2010

Exocelina maculata (Sharp 1882)

Exocelina madangensis (Balke 2001)

Exocelina manfredi (Balke 1998)

Exocelina marinae (Shaverdo, Sagata & Balke 2005)

Exocelina me (Balke 1998)

Exocelina melanaria (Sharp 1882)

Exocelina messeri (Balke 1999)

Exocelina miriae (Balke 1998)

Exocelina monae (Balke 1998)

Exocelina monteithi Wewalka, Balke & Hendrich 2010

Exocelina munaso (Shaverdo, Sagata & Balke 2005)

Exocelina nielsi Wewalka, Balke & Hendrich 2010

Exocelina niklasi Wewalka, Balke & Hendrich 2010

Exocelina nomax (J. Balfour-Browne 1939)

Exocelina novaecaledoniae (J. Balfour-Browne 1939)

Exocelina ouin Wewalka, Balke & Hendrich 2010

Exocelina parvula (Boisduval 1835)

Exocelina patepensis (Balke 1998)

Exocelina perfecta (Sharp 1882)

Exocelina poellabauerae Wewalka, Balke & Hendrich 2010

Exocelina polita (Sharp 1882)

Exocelina punctipennis (Lea 1899)

Exocelina rasilis (Lea 1899)

Exocelina rasjadi Watts & Humphreys 2009

Exocelina remyi Wewalka, Balke & Hendrich 2010

Exocelina rivulus (Balke 1998)

Exocelina rotteri Wewalka, Balke & Hendrich 2010

Exocelina rufa (Balke 1998)

Exocelina sanctimontis (Balke 1998)

Exocelina schoelleri Wewalka, Balke & Hendrich 2010

Exocelina shizong (Balke & Bergsten 2003)

Exocelina simoni Wewalka, Balke & Hendrich 2010

Exocelina simplex (Clark 1863)

Exocelina staneki Wewalka, Balke & Hendrich 2010

Exocelina subjecta (Sharp 1882)

Exocelina takime (Balke 1998)

Exocelina talaki (Balke 1998)

Exocelina tarmluensis (Balke 1998)

Exocelina ullrichi (Balke 1998)

Exocelina vladimiri (Shaverdo, Sagata & Balke 2005)

Acilius

Aethionectes

Africodytes

Africophilus

Agabinus

Agabus

Agaporomorphus

Aglymbus

Allomatus

Allopachria

Amarodytes

Andex

Anginopachria

Anisomeria

Bidessodes

Canthyporus

Carabdytes

Carabhydrus

Coelhydrus

Colymbetes

Coptotomus

Crinodessus

Deronectes

Derovatellus

Dimitshydrus

Ereboporus

Eretes

Fontidessus

Graphoderus

Heroceras

Herophydrus

Heterosternuta

Hoperius

Huxelhydrus

Hydaticus

Hyderodes

Hydrodessus

Hydrodytes

Hydroglyphus

Hydronebrius

Hydroporus

Hydrotrupes

Hygrotus

Hyphoporus

Hyphydrus

Hypodessus

Iberoporus

Ilybiosoma

Ilybius

Japanolaccophilus

Kuschelydrus

Laccodytes

Lacconectus

Laccophilus

Laccornellus

Laccornis

Laccosternus

Leiodytes

Limbodessus

Madaglymbus

Megadytes

Melanodytes

Metaporus

Methles

Microdessus

Microhydrodytes

Napodytes

Nebrioporus

Neobidessus

Neoclypeodytes

Neoscutopterus

Neptosternus

Notaticus

Onychohydrus

Oreodytes

Pachynectes

Paroster

Peschetius

Philaccolilus

Phreatodessus

Platambus

Platydytes

Platynectes

Primospes

Rhithrodytes

Rugosus

Sandracottus

Scarodytes

Senilites

Siamoporus

Siettitia

Spencerhydrus

Sternopriscus

Stictonectes

Tepuidessus

Trichonectes

Trogloguignotus

Typhlodessus

Yola

Yolina

78 additional siblings truncated for brevity.

See the resource file for a full list.

Life

Cellular Organisms

Eukaryota

Opisthokonta

Metazoa

Bilateria

Protostomia

Ecdysozoa

Panarthropoda

Arthropoda

Pancrustacea

Hexapoda

Insecta

Pterygota

Neoptera

Endopterygota

Coleoptera

Adephaga

Dytiscidae

Exocelina (this page)

Exocelina abdita (Balke, Watts, Cooper, Humphreys & Vogler 2004)

Exocelina aipo (Balke 1998)

Exocelina aipomek (Balke 1998)

Exocelina allerbergeri Wewalka, Balke & Hendrich 2010

Exocelina ascendens (Balke 1998)

Exocelina astrophallus (Balke 1998)

Exocelina atowaso (Shaverdo, Sagata & Balke 2005)

Exocelina atra (Sharp 1882)

Exocelina atrata (J. Balfour-Browne 1939)

Exocelina atripennis (J. Balfour-Browne 1939)

Exocelina aubei (Montrouzier 1860)

Exocelina australiae (Clark 1863)

Exocelina australis (Clark 1863)

Exocelina bacchusi (Balke 1998)

Exocelina bagus (Balke & Hendrich ex Balke 2001)

Exocelina barbarae Wewalka, Balke & Hendrich 2010

Exocelina bimaculata (Perroud & Montrouzier 1864)

Exocelina boulevardi (Watts 1978)

Exocelina broschii (Balke 1998)

Exocelina brownei (Guignot 1942)

Exocelina brunoi Wewalka, Balke & Hendrich 2010

Exocelina burwelli Wewalka, Balke & Hendrich 2010

Exocelina casuarina (Balke & Hendrich 1998)

Exocelina charlottae Wewalka, Balke & Hendrich 2010

Exocelina commatifera (Heller 1916)

Exocelina creuxorum Wewalka, Balke & Hendrich 2010

Exocelina damantiensis (Balke 1998)

Exocelina danae (Balke 1998)

Exocelina desii (Balke 1999)

Exocelina elongatula (W. J. Mac Leay 1871)

Exocelina erteldi (Balke 1998)

Exocelina ferruginea (Sharp 1882)

Exocelina feryi Wewalka, Balke & Hendrich 2010

Exocelina flammi Wewalka, Balke & Hendrich 2010

Exocelina fume (Balke 1998)

Exocelina gapa (Watts 1978)

Exocelina gaulorum Wewalka, Balke & Hendrich 2010

Exocelina gelima Wewalka, Balke & Hendrich 2010

Exocelina glypta (Guignot 1955)

Exocelina gracilis (Sharp 1882)

Exocelina heidiae (Balke 1998)

Exocelina hintelmannae (Shaverdo, Sagata & Balke 2005)

Exocelina inexspectata Wewalka, Balke & Hendrich 2010

Exocelina interrupta (Perroud & Montrouzier 1864)

Exocelina jaseminae (Balke 1998)

Exocelina jeannae Wewalka, Balke & Hendrich 2010

Exocelina kainantuensis (Balke 2001)

Exocelina karmurensis (Balke 1998)

Exocelina ketembang (Balke 1998)

Exocelina koghis Wewalka, Balke & Hendrich 2010

Exocelina kolleri Wewalka, Balke & Hendrich 2010

Exocelina larsoni (Balke 1998)

Exocelina leae Wewalka, Balke & Hendrich 2010

Exocelina lilianae Wewalka, Balke & Hendrich 2010

Exocelina maculata (Sharp 1882)

Exocelina madangensis (Balke 2001)

Exocelina manfredi (Balke 1998)

Exocelina marinae (Shaverdo, Sagata & Balke 2005)

Exocelina me (Balke 1998)

Exocelina melanaria (Sharp 1882)

Exocelina messeri (Balke 1999)

Exocelina miriae (Balke 1998)

Exocelina monae (Balke 1998)

Exocelina monteithi Wewalka, Balke & Hendrich 2010

Exocelina munaso (Shaverdo, Sagata & Balke 2005)

Exocelina nielsi Wewalka, Balke & Hendrich 2010

Exocelina niklasi Wewalka, Balke & Hendrich 2010

Exocelina nomax (J. Balfour-Browne 1939)

Exocelina novaecaledoniae (J. Balfour-Browne 1939)

Exocelina ouin Wewalka, Balke & Hendrich 2010

Exocelina parvula (Boisduval 1835)

Exocelina patepensis (Balke 1998)

Exocelina perfecta (Sharp 1882)

Exocelina poellabauerae Wewalka, Balke & Hendrich 2010

Exocelina polita (Sharp 1882)

Exocelina punctipennis (Lea 1899)

Exocelina rasilis (Lea 1899)

Exocelina rasjadi Watts & Humphreys 2009

Exocelina remyi Wewalka, Balke & Hendrich 2010

Exocelina rivulus (Balke 1998)

Exocelina rotteri Wewalka, Balke & Hendrich 2010

Exocelina rufa (Balke 1998)

Exocelina sanctimontis (Balke 1998)

Exocelina schoelleri Wewalka, Balke & Hendrich 2010

Exocelina shizong (Balke & Bergsten 2003)

Exocelina simoni Wewalka, Balke & Hendrich 2010

Exocelina simplex (Clark 1863)

Exocelina staneki Wewalka, Balke & Hendrich 2010

Exocelina subjecta (Sharp 1882)

Exocelina takime (Balke 1998)

Exocelina talaki (Balke 1998)

Exocelina tarmluensis (Balke 1998)

Exocelina ullrichi (Balke 1998)

Exocelina vladimiri (Shaverdo, Sagata & Balke 2005)

Acilius

Africodytes

Africophilus

Agabinus

Agabus

Agaporomorphus

Aglymbus

Allomatus

Amarodytes

Andex

Anginopachria

Anisomeria

Bidessodes

Canthyporus

Carabhydrus

Coelambus

Coelhydrus

Colymbetes

Coptotomus

Crinodessus

Deronectes

Derovatellus

Dimitshydrus

Fontidessus

Glessaria

Graphoderus

Heroceras

Herophydrus

Heterosternuta

Hoperius

Huxelhydrus

Hydaticus

Hydrodessus

Hydrodytes

Hydroglyphus

Hydroporus

Hydrotrupes

Hygrotus

Hyphoporus

Hyphydrus

Hypodessus

Iberoporus

Ilybiosoma

Japanolaccophilus

Kuschelydrus

Laccodytes

Lacconectus

Laccophilus

Laccornellus

Laccornis

Laccosternus

Leiodytes

Liadroporus

Liadyxianus

Limbodessus

Madaglymbus

Megadytes

Melanodytes

Mesoderus

Mesodytes

Metaporus

Microdessus

Microhydrodytes

Napodytes

Nebrioporus

Neobidessus

Neoclypeodytes

Neoscutopterus

Neptosternus

Notaticus

Onychohydrus

Oreodytes

Pachynectes

Palaeodytes

Paroster

Peschetius

Philaccolilus

Phreatodessus

Platambus

Platydytes

Platynectes

Primospes

Procoelambus

Rhithrodytes

Rugosus

Sandracottus

Scarodytes

Senilites

Siamoporus

Siettitia

Sinoporus

Spencerhydrus

Sternopriscus

Stictonectes

Tepuidessus

Trichonectes

Trogloguignotus

Typhlodessus

Yola

Yolina

93 additional siblings truncated for brevity.

See the resource file for a full list.

Arthropoda

Insecta

Coleoptera

Dytiscidae

Exocelina Broun 1886 (this page)

Exocelina abdita (Balke, Watts, Cooper, Humphreys & Vogler 2004)

Exocelina aipo (Balke 1998)

Exocelina aipomek (Balke 1998)

Exocelina allerbergeri Wewalka, Balke & Hendrich 2010

Exocelina ascendens (Balke 1998)

Exocelina astrophallus (Balke 1998)

Exocelina atowaso (Shaverdo, Sagata & Balke 2005)

Exocelina atra (Sharp 1882)

Exocelina atrata (J. Balfour-Browne 1939)

Exocelina atripennis (J. Balfour-Browne 1939)

Exocelina aubei (Montrouzier 1860)

Exocelina australiae (Clark 1863)

Exocelina australis (Clark 1863)

Exocelina bacchusi (Balke 1998)

Exocelina bagus (Balke & Hendrich 2001)

Exocelina barbarae Wewalka, Balke & Hendrich 2010

Exocelina bimaculata (Perroud & Montrouzier 1864)

Exocelina boulevardi (Watts 1978)

Exocelina broschii (Balke 1998)

Exocelina brownei (Guignot 1942)

Exocelina brunoi Wewalka, Balke & Hendrich 2010

Exocelina burwelli Wewalka, Balke & Hendrich 2010

Exocelina casuarina (Balke & Hendrich 1998)

Exocelina charlottae Wewalka, Balke & Hendrich 2010

Exocelina commatifera (Heller 1916)

Exocelina creuxorum Wewalka, Balke & Hendrich 2010

Exocelina damantiensis (Balke 1998)

Exocelina danae (Balke 1998)

Exocelina desii (Balke 1999)

Exocelina elongatula (W. J. Mac Leay 1871)

Exocelina erteldi (Balke 1998)

Exocelina ferruginea (Sharp 1882)

Exocelina feryi Wewalka, Balke & Hendrich 2010

Exocelina flammi Wewalka, Balke & Hendrich 2010

Exocelina fume (Balke 1998)

Exocelina gapa (Watts 1978)

Exocelina gaulorum Wewalka, Balke & Hendrich 2010

Exocelina gelima Wewalka, Balke & Hendrich 2010

Exocelina glypta (Guignot 1955)

Exocelina gracilis (Sharp 1882)

Exocelina heidiae (Balke 1998)

Exocelina hintelmannae (Shaverdo, Sagata & Balke 2005)

Exocelina inexspectata Wewalka, Balke & Hendrich 2010

Exocelina interrupta (Perroud & Montrouzier 1864)

Exocelina jaseminae (Balke 1998)

Exocelina jeannae Wewalka, Balke & Hendrich 2010

Exocelina kainantuensis (Balke 2001)

Exocelina karmurensis (Balke 1998)

Exocelina ketembang (Balke 1998)

Exocelina koghis Wewalka, Balke & Hendrich 2010

Exocelina kolleri Wewalka, Balke & Hendrich 2010

Exocelina larsoni (Balke 1998)

Exocelina leae Wewalka, Balke & Hendrich 2010

Exocelina lilianae Wewalka, Balke & Hendrich 2010

Exocelina maculata (Sharp 1882)

Exocelina madangensis (Balke 2001)

Exocelina manfredi (Balke 1998)

Exocelina marinae (Shaverdo, Sagata & Balke 2005)

Exocelina me (Balke 1998)

Exocelina melanaria (Sharp 1882)

Exocelina messeri (Balke 1999)

Exocelina miriae (Balke 1998)

Exocelina monae (Balke 1998)

Exocelina monteithi Wewalka, Balke & Hendrich 2010

Exocelina munaso (Shaverdo, Sagata & Balke 2005)

Exocelina nehoue Balke, Hájek, Hendrich & Wewalka 2014

Exocelina nielsi Wewalka, Balke & Hendrich 2010

Exocelina niklasi Wewalka, Balke & Hendrich 2010

Exocelina nomax (J. Balfour-Browne 1939)

Exocelina novaecaledoniae (J. Balfour-Browne 1939)

Exocelina ouin Wewalka, Balke & Hendrich 2010

Exocelina parvula (Boisduval 1835)

Exocelina patepensis (Balke 1998)

Exocelina perfecta (Sharp 1882)

Exocelina poellabauerae Wewalka, Balke & Hendrich 2010

Exocelina polita (Sharp 1882)

Exocelina punctipennis (Lea 1899)

Exocelina rasilis (Lea 1899)

Exocelina rasjadi Watts & Humphreys 2009

Exocelina remyi Wewalka, Balke & Hendrich 2010

Exocelina rivulus (Balke 1998)

Exocelina rotteri Wewalka, Balke & Hendrich 2010

Exocelina rufa (Balke 1998)

Exocelina sanctimontis (Balke 1998)

Exocelina schoelleri Wewalka, Balke & Hendrich 2010

Exocelina shizong (Balke & Bergsten 2003)

Exocelina simoni Wewalka, Balke & Hendrich 2010

Exocelina simplex (Clark 1863)

Exocelina staneki Wewalka, Balke & Hendrich 2010

Exocelina subjecta (Sharp 1882)

Exocelina takime (Balke 1998)

Exocelina talaki (Balke 1998)

Exocelina tarmluensis (Balke 1998)

Exocelina ullrichi (Balke 1998)

Exocelina vladimiri (Shaverdo, Sagata & Balke 2005)

Africodytes Biström 1988

Africophilus Guignot 1948

Agabetes Crotch 1873

Agabinus Crotch 1873

Agabus Leach 1817

Agametrus Sharp 1882

Agnoshydrus Biström, Nilsson & Wewalka 1997

Andex Sharp 1882

Angaragabus Ponomarenko 1963

Anisomeria Brinck 1943

Anodocheilus Babington 1841

Bidessodes Régimbart 1900

Bidessonotus Régimbart 1895

Bidessus Sharp 1882

Borneodessus Balke, Hendrich, Mazzoldi & Biström (2002)

Brachyvatus Zimmermann 1919

Celina Aubé 1837

Clypeodytes Régimbart 1894

Coelambus Thomson 1860

Coelhydrus Sharp 1882

Comaldessus Spangler & Barr 1995

Copelatus Erichson 1832

Coptotomus Say 1830

Crinodessus K. B. Miller 1997

Cybister Curtis 1827

Deronectes Sharp 1882

Derovatellus Sharp 1882

Desmopachria Babington 1841

Dytiscus Linnaeus 1758

Eretes Laporte 1832

Glareadessus Wewalka & Biström 1998

Graphoderus Dejean 1833

Graptodytes Seidlitz 1887

Haideoporus Young & Longley 1976

Heroceras Guignot 1950

Herophydrus Sharp 1880

Heterosternuta Strand 1935

Hoperius Fall 1927

Hovahydrus Biström 1982

Hyderodes Hope 1838

Hydrocolus Roughley & Larson 2000

Hydrodessus J. Balfour-Browne 1953

Hydrodytes K. B. Miller (2001)

Hydroglyphus Motschulsky 1853

Hydrotrupes Sharp 1882

Hydrovatus Motschulsky 1853

Hygrotus Stephens 1828

Hyphydrus Illiger 1802

Iberoporus Castro & Delgado 2001

Ilybiosoma Crotch 1873

Ilybius Erichson 1832

Laccodytes Régimbart 1895

Lacconectus Motschulsky 1855

Laccophilus Leach 1815

Laccornis Gozis 1914

Leuronectes Sharp 1882

Liodessus Guignot 1939

Lioporeus Guignot 1950

Liopterus Dejean 1833

Matus Aubé 1836

Megadytes Sharp 1882

Megaporus Brinck 1943

Meladema Laporte 1835

Melanodytes Seidlitz 1887

Metaporus Guignot 1945

Methles Sharp 1882

Microdytes J. Balfour-Browne 1946

Microhydrodytes K. B. Miller (2002)

Miodytiscus Wickham 1911

Nebrioporus Régimbart 1906

Neobidessus Young 1967

Neoclypeodytes Young 1967

Neoporus Guignot 1931

Neoscutopterus Balfour-Browne 1943

Notaticus Zimmermann 1928

Oreodytes Seidlitz 1887

Pachydrus Sharp 1882

Pachynectes Régimbart 1903

Palaeogyrinus Schlechtendal 1894

Paroster Sharp 1882

Philaccolus Guignot 1937

Platambus C. G. Thomson 1859

Platydytes Biström 1988

Porhydrus Guignot 1945

Procoelambus Théobald 1937

Rhithrodytes Bameul 1989

Sanfilippodytes Franciscolo 1979

Scarodytes Des Gozis 1914

Senilites Brinck 1948

Siamoporus Spangler 1996

Siettitia Abeille de Perrin 1904

Stictonectes Brinck 1943

Stictotarsus A. Zimmermann 1919

Suphrodytes Des Gozis 1914

Tepuidessus Spangler 1981

Thermonectus Dejean 1833

Tiporus Watts 1985

Typhlodessus Brancucci 1985

Uvarus Guignot 1939

Yola Gozis 1886

105 additional siblings truncated for brevity.

See the resource file for a full list.

Bilateria

Protostomia

Ecdysozoa

Arthropoda

Hexapoda

Insecta

Pterygota

Neoptera

Holometabola

Coleoptera Linnaeus 1758

Adephaga Schellenberg 1806

Dytiscidae Leach 1815

Copelatinae Branden 1885

Copelatini Branden 1885

Exocelina Broun 1886 (this page)

Exocelina abdita (Balke, Watts, Cooper, Humphreys & Vogler 2004)

Exocelina adelbertensis Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina aipo (Balke 1998)

Exocelina aipomek (Balke 1998)

Exocelina akameku Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina alexanderi Shaverdo, Hendrich & Balke 2012

Exocelina allerbergeri Wewalka, Balke & Hendrich 2010

Exocelina amabilis Shaverdo & Balke ex Shaverdo et al. 2020

Exocelina ambua Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina andakombensis Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina anggiensis Shaverdo, Hendrich & Balke 2012

Exocelina apistefti Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina arfakensis Shaverdo, Hendrich & Balke 2012

Exocelina ascendens (Balke 1998)

Exocelina aseki Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina astrophallus (Balke 1998)

Exocelina athesphati Shaverdo, Surbakti, Sumoked & Balke 2020

Exocelina atowaso (Shaverdo, Sagata & Balke 2005)

Exocelina atra (Sharp 1882)

Exocelina atrata (J. Balfour-Browne 1939)

Exocelina atripennis (J. Balfour-Browne 1939)

Exocelina aubei (Montrouzier 1860)

Exocelina australiae (Clark 1863)

Exocelina australis (Clark 1863)

Exocelina bacchus Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina bagus (Balke & Hendrich ex Balke 2001)

Exocelina baliem Shaverdo, Hendrich & Balke 2013

Exocelina barbarae Wewalka, Balke & Hendrich 2010

Exocelina bewani Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina bewaniensis Shaverdo, Menufandu & Balke 2014

Exocelina bifida Shaverdo, Hendrich & Balke 2012

Exocelina bimaculata (Perroud & Montrouzier 1864)

Exocelina bismarckensis Shaverdo & Balke 2014

Exocelina boulevardi (Watts 1978)

Exocelina brahminensis Shaverdo, Hendrich & Balke 2012

Exocelina brazza Shaverdo & Balke ex Shaverdo et al. 2020

Exocelina broschii (Balke 1998)

Exocelina brownei (Guignot 1942)

Exocelina brunoi Wewalka, Balke & Hendrich 2010

Exocelina bundiensis Shaverdo, Hendrich & Balke 2012

Exocelina burwelli Wewalka, Balke & Hendrich 2010

Exocelina casuarina (Balke & Hendrich 1998)

Exocelina charlottae Wewalka, Balke & Hendrich 2010

Exocelina cheesmanae (J. Balfour-Browne 1939)

Exocelina commatifera (Heller 1916)

Exocelina craterensis Shaverdo & Balke 2014

Exocelina creuxorum Wewalka, Balke & Hendrich 2010

Exocelina cyclops Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina damantiensis (Balke 1998)

Exocelina danae (Balke 1998)

Exocelina desii (Balke 1999)

Exocelina edeltraudae Shaverdo, Hendrich & Balke 2012

Exocelina ekari Shaverdo, Hendrich & Balke 2012

Exocelina ekpliktiki Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina elongatula (W. J. MacLeay 1871)

Exocelina eme Shaverdo, Hendrich & Balke 2012

Exocelina erteldi (Balke 1998)

Exocelina evelyncheesmanae Shaverdo, Hendrich & Balke 2012

Exocelina ferruginea (Sharp 1882)

Exocelina feryi Wewalka, Balke & Hendrich 2010

Exocelina flammi Wewalka, Balke & Hendrich 2010

Exocelina foja Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina fume (Balke 1998)

Exocelina gapa (Watts 1978)

Exocelina garana Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina gaulorum Wewalka, Balke & Hendrich 2010

Exocelina gelima Wewalka, Balke & Hendrich 2010

Exocelina glypta (Guignot 1955)

Exocelina gorokaensis Shaverdo & Balke 2014

Exocelina gracilis (Sharp 1882)

Exocelina haia Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina hansferyi Shaverdo, Hendrich & Balke 2012

Exocelina heidiae (Balke 1998)

Exocelina herowana Shaverdo & Balke 2014

Exocelina hintelmannae (Shaverdo, Sagata & Balke 2005)

Exocelina hudsoni Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina ibalimi Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina inexspectata Wewalka, Balke & Hendrich 2010

Exocelina injiensis Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina interrupta (Perroud & Montrouzier 1864)

Exocelina iratoi Shaverdo, Wild, Sumoked & Balke 2017

Exocelina irianensis Shaverdo, Hendrich & Balke 2012

Exocelina jaseminae (Balke 1998)

Exocelina jeannae Wewalka, Balke & Hendrich 2010

Exocelina jimiensis Shaverdo & Balke 2014

Exocelina kabwumensis Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina kailaki Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina kainantuensis (Balke 2001)

Exocelina kakapupu Shaverdo, Hendrich & Balke 2012

Exocelina karmurensis (Balke 1998)

Exocelina keki Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina ketembang (Balke 1998)

Exocelina kinibeli Shaverdo & Balke 2014

Exocelina kisli Shaverdo & Balke 2014

Exocelina knoepfchen Shaverdo, Hendrich & Balke 2012

Exocelina kobau Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina koghis Wewalka, Balke & Hendrich 2010

Exocelina kolleri Wewalka, Balke & Hendrich 2010

Exocelina koroba Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina kowalskii Shaverdo & Balke 2019

Exocelina ksionseki Shaverdo & Balke 2014

Exocelina kumulensis Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina larsoni (Balke 1998)

Exocelina leae Wewalka, Balke & Hendrich 2010

Exocelina lembena Shaverdo & Balke 2014

Exocelina likui Shaverdo, Wild, Sumoked & Balke 2017

Exocelina lilianae Wewalka, Balke & Hendrich 2010

Exocelina maculata (Sharp 1882)

Exocelina manfredi (Balke 1998)

Exocelina manokwariensis Shaverdo, Panjaitan & Balke 2016

Exocelina mantembu Shaverdo & Balke 2014

Exocelina marawaka Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina marinae (Shaverdo, Sagata & Balke 2005)

Exocelina me (Balke 1998)

Exocelina mekilensis Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina melanaria (Sharp 1882)

Exocelina mendiensis Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina menyamya Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina messeri (Balke 1999)

Exocelina mianminensis Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina michaelensis Shaverdo & Balke 2014

Exocelina mimika Shaverdo & Balke ex Shaverdo et al. 2020

Exocelina miriae (Balke 1998)

Exocelina monae (Balke 1998)

Exocelina mondmillensis Shaverdo, Sagata & Balke 2016

Exocelina monteithi Wewalka, Balke & Hendrich 2010

Exocelina morobensis Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina munaso (Shaverdo, Sagata & Balke 2005)

Exocelina nehoue Balke, Hájek, Hendrich & Wewalka 2014

Exocelina nielsi Wewalka, Balke & Hendrich 2010

Exocelina niklasi Wewalka, Balke & Hendrich 2010

Exocelina nomax (J. Balfour-Browne 1939)

Exocelina novaecaledoniae (J. Balfour-Browne 1939)

Exocelina oceai Shaverdo, Hendrich & Balke 2012

Exocelina oiwa Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina okapa Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina okbapensis Shaverdo & Balke ex Shaverdo et al. 2017

Exocelina oksibilensis Shaverdo, Surbakti, Warikar & Balke ex Shaverdo et al. 2019

Exocelina oraia Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina ouin Wewalka, Balke & Hendrich 2010

Exocelina parvula (Boisduval 1835)

Exocelina perfecta (Sharp 1882)

Exocelina pinocchi Shaverdo & Balke 2014

Exocelina piusi Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina poellabauerae Wewalka, Balke & Hendrich 2010

Exocelina polita (Sharp 1882)

Exocelina posmani Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina pseudoastrophallus Shaverdo & Balke 2014

Exocelina pseudobifida Shaverdo & Balke 2014

Exocelina pseudoedeltraudae Shaverdo & Balke 2014

Exocelina pseudoeme Shaverdo & Balke 2014

Exocelina pseudofume Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina pseudojaseminae Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina pseudomarinae Shaverdo, Sagata & Balke 2016

Exocelina pseudopusilla Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina pseudosoppi Shaverdo, Hendrich & Balke 2012

Exocelina pui Shaverdo, Wild, Sumoked & Balke 2017

Exocelina pulchella Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina pulukensis Shaverdo, Wild, Sumoked & Balke 2017

Exocelina punctipennis (Lea 1899)

Exocelina pusilla Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina ransikiensis Shaverdo, Panjaitan & Balke 2016

Exocelina rasilis (Lea 1899)

Exocelina rasjadi Watts & Humphreys 2009

Exocelina remyi Wewalka, Balke & Hendrich 2010

Exocelina riberai Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina rotteri Wewalka, Balke & Hendrich 2010

Exocelina rufa (Balke 1998)

Exocelina saltusholmesensis Watts, Hendrich & Balke 2016

Exocelina saudaunensis Shaverdo & Balke 2014

Exocelina schoelleri Wewalka, Balke & Hendrich 2010

Exocelina shizong (Balke & Bergsten 2003)

Exocelina sima Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina simbaiarea Shaverdo & Balke 2014

Exocelina simbaiensis Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina simbaijimi Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina simoni Wewalka, Balke & Hendrich 2010

Exocelina simplex (Clark 1863)

Exocelina skalei Shaverdo & Balke 2014

Exocelina soppi Shaverdo, Hendrich & Balke 2012

Exocelina staneki Wewalka, Balke & Hendrich 2010

Exocelina subjecta (Sharp 1882)

Exocelina sugayai Balke & Ribera 2020

Exocelina sumokedi Shaverdo & Balke ex Shaverdo et al. 2018

Exocelina tabubilensis Shaverdo & Balke 2014

Exocelina takime (Balke 1998)

Exocelina talaki (Balke 1998)

Exocelina tariensis Shaverdo & Balke 2014

Exocelina tekadu Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina tomhansi Shaverdo, Wild, Sumoked & Balke 2017

Exocelina tsinga Shaverdo, Surbakti, Sumoked & Balke 2020

Exocelina ullrichi (Balke 1998)

Exocelina unipo Shaverdo, Hendrich & Balke 2012

Exocelina utowaensis Shaverdo, Hendrich & Balke 2012

Exocelina varirata Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina vladimiri (Shaverdo, Sagata & Balke 2005)

Exocelina vovai Shaverdo & Balke 2014

Exocelina waaf Shaverdo, Surbakti & Balke ex Shaverdo, Surbakti, Sumoked & Balke 2021

Exocelina waigeoensis Shaverdo, Hendrich & Balke 2012

Exocelina wannangensis Shaverdo & Balke 2014

Exocelina warahulenensis Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina warasera Shaverdo & Balke ex Shaverdo et al. 2019

Exocelina wareaga Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina weylandensis Shaverdo, Hendrich & Balke 2012

Exocelina wigodukensis Shaverdo, Wild, Sumoked & Balke 2017

Exocelina woitapensis Shaverdo & Balke ex Shaverdo et al. 2016

Exocelina wondiwoiensis Shaverdo, Hendrich & Balke 2012

Exocelina yoginofi Shaverdo & Balke ex Shaverdo et al. 2018

Agaporomorphus Zimmermann 1921

Aglymbus Sharp 1880

Capelatus Turner & Bilton 2015

Copelatus Erichson 1832

Lacconectus Motschulsky 1855

Liopterus Dejean 1833

Madaglymbus Shaverdo & Balke ex Shaverdo, Monaghan, Lees, Ranaivosolo & Balke 2008

root

cellular organisms

Eukaryota

Opisthokonta

Metazoa

Eumetazoa

Bilateria

Protostomia

Ecdysozoa

Panarthropoda

Arthropoda

Mandibulata

Pancrustacea

Hexapoda

Insecta

Dicondylia

Pterygota

Neoptera

Holometabola

Coleoptera

Adephaga

Dytiscoidea

Dytiscidae

Copelatinae

Exocelina (this page)

Exocelina abdita

Exocelina aipomek

Exocelina anggiensis

Exocelina arfakensis

Exocelina astrophallus

Exocelina atowaso

Exocelina atra

Exocelina aubei

Exocelina australiae

Exocelina bacchusi

Exocelina bagus

Exocelina barbarae

Exocelina bimaculata

Exocelina boulevardi

Exocelina brahminensis

Exocelina broschii

Exocelina brownei

Exocelina brunoi

Exocelina bundiensis

Exocelina burwelli

Exocelina charlottae

Exocelina commatifera

Exocelina creuxorum

Exocelina damantiensis

Exocelina desii

Exocelina edeltraudae

Exocelina elongatula

Exocelina evelyncheesmanae

Exocelina ferruginea

Exocelina feryi

Exocelina flammi

Exocelina fume

Exocelina gapa

Exocelina gaulorum

Exocelina gelima

Exocelina hintelmannae

Exocelina interrupta

Exocelina irianensis

Exocelina jaseminae

Exocelina kakapupu

Exocelina knoepfchen

Exocelina larsoni

Exocelina leae

Exocelina maculata

Exocelina madangensis

Exocelina manokwariensis

Exocelina marinae

Exocelina melanaria

Exocelina miriae

Exocelina monae

Exocelina monteithi

Exocelina munaso

Exocelina nielsi

Exocelina niklasi

Exocelina novaecaledoniae

Exocelina ouin

Exocelina parvula

Exocelina patepensis

Exocelina perfecta

Exocelina poellabauerae

Exocelina pseudobifida

Exocelina punctipennis

Exocelina remyi

Exocelina rotteri

Exocelina sandaunensis

Exocelina shizong

Exocelina simoni

Exocelina simplex

Exocelina soppi

Exocelina staneki

Exocelina subjecta

Exocelina takime

Exocelina ullrichi

Exocelina unipo

Exocelina utowaensis

Exocelina weylandensis

Exocelina wondiwoiensis

unclassified Exocelina

Agaporomorphus

Aglymbus

Capelatus

Copelatus

Lacconectus

Liopterus

Madaglymbus

Papuadytes

unclassified Copelatinae

Biota

Eukaryota

Unikonta

Opisthokonta

Holozoa

Filozoa

Apoikozoa

Animalia

Eumetazoa

Bilateria

Protostomia

Ecdysozoa

Arthropoda

Hexapoda

Insecta

Dicondylia

Pterygota

Metapterygota

Neoptera

Eumetabola

Endopterygota

Coleoptera

Adephaga

Dytiscidae

Copelatinae

Exocelina (this page)

Exocelina abdita

Exocelina adelbertensis

Exocelina aipo

Exocelina aipomek

Exocelina alexanderi

Exocelina allerbergeri

Exocelina ambua

Exocelina andakombensis

Exocelina anggiensis

Exocelina apistefti

Exocelina arfakensis

Exocelina ascendens

Exocelina astrophallus

Exocelina atowaso

Exocelina atra

Exocelina atrata

Exocelina atripennis

Exocelina aubei

Exocelina australiae

Exocelina australis

Exocelina bacchusi

Exocelina bagus

Exocelina baliem

Exocelina barbarae

Exocelina bewani

Exocelina bewaniensis

Exocelina bifida

Exocelina bimaculata

Exocelina bismarckensis

Exocelina boulevardi

Exocelina brahminensis

Exocelina broschii

Exocelina brownei

Exocelina brunoi

Exocelina bundiensis

Exocelina burwelli

Exocelina casuarina

Exocelina charlottae

Exocelina commatifera

Exocelina craterensis

Exocelina creuxorum

Exocelina cyclops

Exocelina damantiensis

Exocelina danae

Exocelina desii

Exocelina edeltraudae

Exocelina ekari

Exocelina ekpliktiki

Exocelina elongatula

Exocelina eme

Exocelina erteldi

Exocelina evelyncheesmanae

Exocelina ferruginea

Exocelina feryi

Exocelina flammi

Exocelina foja

Exocelina fume

Exocelina gapa

Exocelina garaina

Exocelina garana

Exocelina gaulorum

Exocelina gelima

Exocelina glypta

Exocelina gorokaensis

Exocelina gracilis

Exocelina hansferyi

Exocelina heidiae

Exocelina herowana

Exocelina hintelmannae

Exocelina hudsoni

Exocelina ibalimi

Exocelina inexspectata

Exocelina injiensis

Exocelina interrupta

Exocelina iratoi

Exocelina irianensis

Exocelina jaseminae

Exocelina jeannae

Exocelina jimiensis

Exocelina kabwumensis

Exocelina kainantuensis

Exocelina kakapupu

Exocelina karmurensis

Exocelina keki

Exocelina ketembang

Exocelina kinibeli

Exocelina kisli

Exocelina knoepfchen

Exocelina koghis

Exocelina kolleri

Exocelina ksionseki

Exocelina kumulensis

Exocelina larsoni

Exocelina leae

Exocelina lembena

Exocelina likui

Exocelina lilianae

Exocelina maculata

Exocelina madangensis

Exocelina manfredi

Exocelina manokwariensis

Exocelina mantembu

Exocelina marawaka

Exocelina marinae

Exocelina me

Exocelina melanaria

Exocelina mendiensis

Exocelina menyamya

Exocelina messeri

Exocelina michaelensis

Exocelina miriae

Exocelina monae

Exocelina mondmillensis

Exocelina monteithi

Exocelina munaso

Exocelina nehoue

Exocelina nielsi

Exocelina niklasi

Exocelina nomax

Exocelina novaecaledoniae

Exocelina oceai

Exocelina okapa

Exocelina oraia

Exocelina ouin

Exocelina parvula

Exocelina patepensis

Exocelina perfecta

Exocelina pinocchio

Exocelina piusi

Exocelina poellabauerae

Exocelina polita

Exocelina posmani

Exocelina pseudoastrophallus

Exocelina pseudobifida

Exocelina pseudoedeltraudae

Exocelina pseudoeme

Exocelina pseudofume

Exocelina pseudomarinae

Exocelina pseudopusilla

Exocelina pseudosoppi

Exocelina pui

Exocelina pulukensis

Exocelina punctipennis

Exocelina pusilla

Exocelina ransikiensis

Exocelina rasilis

Exocelina rasjadi

Exocelina remyi

Exocelina riberai

Exocelina rivulus

Exocelina rotteri

Exocelina rufa

Exocelina saltusholmesensis

Exocelina sanctimontis

Exocelina sandaunensis

Exocelina schoelleri

Exocelina shizong

Exocelina sima

Exocelina simbaiarea

Exocelina simbaiensis

Exocelina simbaijimi

Exocelina simoni

Exocelina simplex

Exocelina skalei

Exocelina soppi

Exocelina staneki

Exocelina subjecta

Exocelina sugayai

Exocelina sumokedi

Exocelina tabubilensis

Exocelina takime

Exocelina talaki

Exocelina tariensis

Exocelina tarmluensis

Exocelina tekadu

Exocelina tomhansi

Exocelina ullrichi

Exocelina unipo

Exocelina utowaensis

Exocelina varirata

Exocelina vladimiri

Exocelina vovai

Exocelina waaf

Exocelina waigeoensis

Exocelina wannangensis

Exocelina wareaga

Exocelina weylandensis

Exocelina wigodukensis

Exocelina woitapensis

Exocelina wondiwoiensis

Exocelina yoginofi

Capelatus

Copelatini

Copelatus