Speciomerus revoili

Speciomerus revoili (Pic) Caryoborus revoili Pic 1902:172.
Pachymerus revoili: Pic 1913:8.
Caryobruchus revoili: Bridwell 1929:155; Udayagiri and
Wadhi 1989:240. Caryobruchus acrocomiae Bridwell 1929:153; Udayagiri
and Wadhi 1989:238. New synonymy.
Type Data. — Type locality: Paraguay. Holotype female, Revoil; deposited in Museum National d'Histoire Naturelle, Paris (MNHP). The type is badly insect damaged, a homotype was therefore deposited in MNHP together with the type. No allotype and no other paratypes. Caryoborus acrocomiae: Holotype, Uruguay, ex seed Acrocomia sp., quarantine Washington , D.C, 9.XII.1921; deposited in the U.S. National Museum, Washington, Distribution. — (Fig. 2). Specimens examined, 52. Argentina (Salta, Misiones), Bolivia (Santa Cruz), Brazil (Mato Grosso, Mato Grosso del Sul, Rio Grande do Sul), Paraguay (Alto Parana, Cordillera, Paraguari, San Pedro), Uruguay, Venezuela ( Apure, Guarico, Barinas).
Diagnosis. — Integument dark reddish-brown, to almost black, some specimens more red than others, appendages and antennae with slightly lighter, more yellowish color, hind femur often with a darker area in the center, fine white to yellowish-white pubescence partly covers integument color. Males 10.3-17.2 mm, females 10.7-16.8 mm in length (pronotum-elytra); width of males 5.6-9.4 mm, females 6.0-8.8 mm, maximum thoracic depth of males 3.8-6.8 mm, females 4.1-7.0 mm. Eyes approximate. Antennal segments 4-10 feebly serrate, segments longer than wide, segment 9 usually longer than segment 4. Disk of pronotum wider than long (length /width ratio mean 0.74). Metafemur usually twice as long as wide (length/width ratio mean 2.05); pecten short with 8-13 denticles, denticle 1 acuminate and longer; prepectenal ridge longer than pecten, with 4-9 usually large acute or edged spines. Metatibia with inner ventral carina not elevated. Pygidium wider than long (length/width ratio mean 0.79).
Males. — Integument. Hirsute, base of each seta in a minute puncture (therefore minutely punctulate
where setae have been removed). Gula, ventral part of gena, apical end of proepimeron (with white setae lining the margin), metacoxa near trochanter, and part of mesal side of metafemur glabrous. Color. Body dark reddish-brown, to almost black, some specimens more red than others, appendages and antennae usually with slightly lighter color, hind femur often with darker area in the center.
Vestiture. Setae often lost on parts of the body, possible due to rubbing. Dorsal and ventral body surfaces with white to yellowish-white, fine (very fine on elytra), recumbent, dense, setae; antenna with minute yellowishwhite setae; elytron surface densely hirsute; protibia and mesotibia with coarse, golden, moderately dense to dense setae apically; metatibia lacking golden setae except a few golden setae inside apex of mucro; ventral ridge of metafemur with sparse setae not covering pecten; tarsomere 1 with moderately dense, golden setae, ventral side of tarsomere 2 with a few coarse, golden setae, ventral side of tarsomere 3 with very dense, coarse, golden setae; py gidium not as densely hirsute as elytron, setae longer and coarser.
Head. Slightly shorter than pronotum; frons and vertex with weak to strong median usually glabrous but sometimes hirsute carina at vertex not interrupted by a weak depression (Fig. 96); ocular sinus vague; on dorsal side eyes approximate; postocular lobe short, with a glabrous carina continuing on gena; antenna clearly shorter than elytra, segments 4-10 serrated, longer than wide, segment 9 usually longer than segment 4, both segments about the same width, or segment 4 slightly wider; gena short; sides of submenrum parallel, not converging triangularly between eyes; on ventral side eyes approximate.
Prothorax. Disk wider than long (length/width ratio mean 0.74); all sides with strong, impressed, marginal line, anterior margin emarginate; angles vaguely pointed, visible in dorsal view; medial dorsal surface flattened to convex, punctate, small punctations (different from the punctations due to loss of setae) evenly distributed over the disk, large punctae unevenly distributed as four clusters; sides depressed, more depressed toward apex, angulate, narrowed to apex from slightly more than middle of side; angles of lateral margin at base produced, angle more than 90°; posterior margin arcuate, produced medially; strong lateral prothoracic carina complete, extending from base to apex. Protarsus 1 shorter and not very wide or swollen
but wider than metatarsus 1 (sexual dimorphism: not wider in females). Prosternum completely separating procoxae, apex of prosternal process narrow or medium width.
Mesothorax and Metathorax. Scutellum small, subquadrate. Elytron with dorsal surface flattened, or sometimes feebly convex, between humerus and mesal margin, depressed laterally and apically; humerus non-scabrous, usually without setae; striae punctate. Metafemur incrassate, maximum width slightly closer to base, constricted toward apex, usually more than twice as long as wide (length/ width ratio mean 2.05), ventral side with a pecten armed with 8-13 denticles, produced, short, straight; denticle 1 large, acuminate, closer to femoral apex than to base; following denticles smaller and decreasing in size; when leg flexed, tibia positioned on lateral side of pecten; prepectenal ridge longer than pecten, with 4-9 usually large acute or edged spines; lateral side with visible small punctures, feebly visible under setae. Metatibia evenly arcuate (angle >90°); three strong ventral carinae, inner carina very strong usually not elevated, wide tibial furrow between inner and middle carinae, middle carina very strong, very close to and fusing with outer carina before mucro, outer carina medium strength, slightly weaker before fusing with middle carinae; lateral, dorsolateral, mesal, and dorsomesal carinae obsolete; mucro long; tibial corona smooth, without spines; spur visible in lateral view, sparsely covered with setae. Mesotarsus 1 shorter, not very wide or swollen but wider than metatarsus 1. Mesosternal process often with a cleft at apex, slanting, projecting to much less than 90° angle from body plane, apex gradually or sharply curved back towards plane of metasternum; metaspinasternum not or only feebly produced.
Abdomen. Sternum 1 about as long as remaining sterna . Py gidium punctulate, convex in lateral view, wider than long (length/width ratio mean 0.79), wider at base than apex, partly hidden by the elytra, elytra cover 30-60% of pygidium length from base, at almost 90° angle to elytron, usually not visible in dorsal view.
Genitalia. Medianlobe (Fig. 137) slender. Ventral valve long with sides forming a triangle with an acute angle. Armature of internal sac without median sclerites; with a transparent, almost colorless, small, median structure, with a cleft and numerous small denticles; and with two very long, slender, basal sclerites, with small denticles at basal end (often hard to see). Lateral lobes (Fig. 138)
confluent, not separated, but with large (for the genus) cleft, apex at about the same level as cutis, with vague median notch, vague fold-like sinus extending from notch to 0.5 length of distance apex to Kingsolver's band.
Females. — Similar to males, but maximum thoracic depth slightly larger. Armature of bursa copulatrix (Fig. 136); one pair of ventral sclerites; one dorsal sclerite. Apodeme of spiculum gastrale normal length.
Specific Records. — Host records are from collection labels, spelled exactly as written, and should be verified. Argentina: Salta, C. Bruch (BR); Misiones, San Ignacio, B. & W. Bade (BR). Bolivia: Santa Cruz, Prov. Andrez Ibanez, Sta. Cruz, Los Olives, 27.11.1989, P. Bettela (FMNH); Santa Cruz, Prov. Andrez Ibanez, 6km S de SC, 10.XII.1989, 30.XII.1989, Bettela (FMNH); Santa Cruz, Province del Sara, II. 1912, XL 1912, Steinbach (CMNH); Santa Cruz, Prov. del Sara, Buena Vista, alt. 1700 ft., IIIV.1925 (ANSP); Ichilo, Buena Vista, alt. 400 m, X.1962 (MSU); Prov. Ichilo, Buena Vista, 61990, P. Bettella (FMNH). Brazil: Mato Grosso, Diamantino, Facienda Sao Joao, alt. 450 m, 6.II.1981, Ekis & Young (CMNH); Mato Grosso del Sul, Corumba, Serra de Urucum, 25.XI.1960, K. Lenko (USNM); Rio Grande do Sul, Chapada (CMNH). Paraguay: Alto Parana, Primavera, 17VIII.1955, E.J. Phillips (BMNH); Cordillera, Emboscada,15.X.1954(CNC); Paraguari, Sapucay, 1903, W. Foster (BMNH); San Pedro, Carumbe, 10-10.11.1973, R. Goldbach (FML); Revoil, holotype (MHNP); in Acrocomia totai Mart., 1951 (USNM). Uruguay: ex seed Acrocomia sp., quarantine Washington , D.C., 9.XII.1921, holotype, and 1 paratype of Caryobruchus acrocomiae (USNM). Venezuela: Apure, Madre Viejo, Rio Apure, en barbas de palma, 21.11.1979, J. Pulido (UCV); Guarico, 59 km E. of Santa Maria de Ipire, from seeds of palm, 15.111.1989 (emerged 30.111.1989), CD. Johnson (CDJ); Barinas, 25 km S. of Barinas, from seed of palm, 19.111.1989, CD. Johnson (CDJ).