Bucculatrix domicola Braun 1963

Bucculatrix domicola new species Face creamy white, tuft ocherous anteriorly and laterally, brown posteriorly ; eye-caps creamy white, scales rarely minutely dark-tipped, antennal stalk annulate with dark brown. Thorax pale ocherous to brownish ocherous, scales rarely dark-tipped, extreme base of fore wing pale when the thorax is pale. Fore wings brownish ocherous, darkest in the costal area between the silvery streaks and on dorsum, especially between the patch of raised scales and tornus. Marks silvery iridescent in both sexes, but sometimes so encroached upon by the brownish scales of the ground color as to be reduced to mere silvery lines, or sometimes ill-defined and not silvery; three parallel and equidistant oblique costal streaks, the first from basal fourth, the second near middle, the third, at two-thirds of costa, meets or nearly meets at an angle in the middle of the wing a similar slender dorsal streak from just before tornus; a few silvery scales precede the black apical spot in which there are a tew raised scales; a large patch of dark brown raised scales on middle of dorsum, extending to fold; often a quadrate patch of paler ground color basad of the patch of raised scales; a few raised scales at tornus; a faint line of dark-tipped scales extends from the apical spot along termen near the base of the apical cilia; a second line of dark browntipped scales curves around apex and extends to tornus, lying closely parallel to the first; cilia varying from ocherous to pale fuscous, darker at tornus. Hind wings and cilia fuscous, darker in males. Legs ocherous. fuscous outwardly, tarsal segments dark-tipped. Abdomen grayish ocherous below, dark fuscous above.
Alar expanse 7 to 7.5 mm.
Male genitalia (tig. 190). Harpes typical of the section, nearly attaining the socii, setose outwardly and sparsely setose inwardly, terminating in a small pointed process, a small basal process present; socii short, broad and widely separated, the sinus between them shallow ; aedeagus gradually tapering to the narrow acute tip; vinculum a very narrow band. Scale sac large, scales slender.
Female genitalia ( hg<. 191, 191a). Fringing scales of the posterior margins of segment 7 long and slender, lobes of sternite of 7 sculptured ; on intersegmental membrane near posterior ventral margin of 7. an arc of specialized scales of moderate length ; posterior to ostium (but anterior to the arc) a median dense patch of small specialized scales ; on the anterior margin of tergite of 8, a mass oi minute scales; margin of ostium sclerotized. with two outwardly directed acutely pointed processes; signum ribs with spines grading from long acute to minute short.
Type. — 3. Cincinnati, Ohio, rearing record B.22P>. with cocoon, imago August 19. 1955 (A. F. Braun) [A.F.B.Coll.].
Allotype. — 9. Cincinnati. Ohio, rearing record B.2219, with cocoon, imago August 19. 1955 (A. F. Braun) [A.F.B.Coll.].
Paratypes. — 2 3. Cincinnati. Ohio. March 2; 2 3 , 4 9. Cincinnati, Ohio, rearing record B.2219, imagoes April 23 ; 1 9, with cocoon, imago June 29; 4 6.2 9, rearing record B.2219. imagoes July 4. July 5; 12 3.6 9. with cocoons, rearing record B.2219. imagoes August 10 to September 2 (A. F. Braun | [A.F.B.Coll.]; 1 3 . New Lisbon, New Jersey. June 2. 1940. emerging from cocoon on underside of a leaf ( E. P. Darlington) [A.X.S.P.] ; 1 3. Monroe County. New York. May 19 ( C. P. Kimball) [C.P.K.Coll.].
In addition to the specimens listed above, one cocoon was collected August 11. near West Union, Adams County', Ohio.
The specimens of the type series from Cincinnati emerged from cocoons spun by larvae feeding on leaves high up in trees of Quercus palustris Muenchh., and Quercus shumardii Buckl. The egg is deposited on the upper side of the leaf against the midrib or a lateral vein. The mine (figs. 54a, 54b) is short, somewhat contorted, thus differing" from that of B. packardella often occurring on the same trees with this species. The larva commonly, in the June and later summer generations, descends to the ground by a silken thread, spinning its cocoon on any convenient surface. The cocoon (fig. 54c) is pale whitish stramineous, slender, elongate, with six or seven prominent ridges.
Three generations a year have been observed and specimens reared at Cincinnati ; moths emerge from overwintering pupae from early March to late April; the larvae of a second generation become full-fed and pupate about mid-June, the imagoes emerging the last of June and early July; a third generation pupates early in August, imagoes emerging in August and early September ; larvae from eggs laid by this generation become full-fed and pupate in late September or October.
Buccidatrix domicola is allied to both B. trifasciella and B. quinquenotella, and males may be mistaken for small males of trifasciella; from qiiinquenotella it is at once distinguished by the less conspicuous silvery streaks and the absence of a silvery spot basad of the raised scales. This species often occurs together with packardella ; worn or pale specimens with obscure markings may be distinguished from that species by the darker hind wings and dark fuscous upper side of the abdomen.
The more slender tapering aedeagus of the male, and the mass of minute specialized scales on the dorsal anterior margin of segment 8 in the female are the most distinctive characters of the genitalia. From all other oak-feeding species it is separated by the slender few-ribbed cocoon.

Bucculatrix domicola is a moth in the family Bucculatricidae. It is found in North America, where it has been recorded from Ohio and New Jersey. It was first described in 1963 by Annette Frances Braun.

The wingspan is 7.0-7.5 mm.

The larvae feed on Quercus palustris and Quercus shumardii. They mine the leaves of their host plants.