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Bishop Pine

Pinus muricata D. Don

Comments

provided by eFloras
The several varieties described for Pinus muricata reflect the high variability in leaf characters and in degree of elaboration of apophysis and umbo in this species. The extremes can sometimes occur together.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 2 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Description

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Trees to 24m; trunk to 0.9m diam., straight to contorted; crown becoming rounded, flattened, or irregular. Bark dark gray, deeply furrowed, ridges long, scaly-plated. Branches spreading-ascending, often contorted; twigs stout to slender, orange-brown, aging darker brown, rough. Buds ovoid-cylindric, dark brown, 1--2.5cm, resinous. Leaves 2 per fascicle, spreading to upcurved, persisting 2--3 years, 8--15cm ´ (1.2--)1.5(--2)mm, slightly twisted, dark yellow-green, all surfaces with stomatal lines, margins strongly serrulate, apex abruptly conic-acute; sheath to 1.5cm, base persistent. Pollen cones ellipsoid, to 5mm, orange. Seed cones maturing in 3 years, serotinous, long-persistent, mostly in whorls, mostly asymmetric, lanceoloid-ovoid before opening, curved-ovoid when open, 4--9cm, glossy bright to pale red-brown, sessile or on stalks to 1cm, mostly downcurved, scales with deep red-brown border distally on adaxial surface; apophyses much thickened, the abaxial ones progressively more angulately dome-shaped toward base of cone; umbo central, a stout-based, curved claw. Seeds obliquely ellipsoid; body 6--7mm, dark brown to near black; wing 15--20mm. 2 n =24.
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cc-by-nc-sa-3.0
copyright
Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 2 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of North America @ eFloras.org
editor
Flora of North America Editorial Committee
project
eFloras.org
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Habitat & Distribution

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Dry ridges to coastal, windshorn forests, often in or around bogs; of conservation concern; 0--300m; Calif.; Mexico in Baja California.
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cc-by-nc-sa-3.0
copyright
Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 2 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of North America @ eFloras.org
editor
Flora of North America Editorial Committee
project
eFloras.org
original
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eFloras

Synonym

provided by eFloras
Pinus muricata var. borealis Axelrod; P. muricata var. cedrosensis J.T. Howell; P. muricata var. stantonii Axelrod; P. radiata var. binata (Engelmann) Brewer & S.Watson; P. remorata H.Mason
license
cc-by-nc-sa-3.0
copyright
Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 2 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of North America @ eFloras.org
editor
Flora of North America Editorial Committee
project
eFloras.org
original
visit source
partner site
eFloras

Common Names

provided by Fire Effects Information System Plants
More info for the terms: cone, swamp, tree

Bishop pine
Bishop's pine
prickle-cone pine
California swamp pine


TAXONOMY:
The scientific name of Bishop pine is Pinus muricata D. Don [8,22,35].

Bishop pine rarely hybridizes with Monterey pine (P. radiata); timing of
cone opening usually differs in the two species [8,44].


LIFE FORM:
Tree

FEDERAL LEGAL STATUS:
No special status

OTHER STATUS:
NO-ENTRY





DISTRIBUTION AND OCCURRENCE
SPECIES: Pinus muricata
GENERAL DISTRIBUTION:
Bishop pine occurs in disjunct coastal populations from Curry
County, Oregon, into California from Humboldt County south
to Santa Barbara County. It is also found on Santa Cruz and
Santa Rosa islands, and in Baja California, Mexico [8,21,22,28,43].
license
cc-publicdomain
bibliographic citation
Cope, Amy B. 1993. Pinus muricata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Distribution

provided by Fire Effects Information System Plants
Bishop pine occurs in disjunct coastal populations from Curry
County, Oregon, into California from Humboldt County south
to Santa Barbara County. It is also found on Santa Cruz and
Santa Rosa islands, and in Baja California, Mexico [8,21,22,28,43].



Distribution of Bishop pine. Map courtesy of USDA, NRCS. 2018. The PLANTS Database.
National Plant Data Team, Greensboro, NC [2018, June 19] [43].

license
cc-publicdomain
bibliographic citation
Cope, Amy B. 1993. Pinus muricata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Fire Management Considerations

provided by Fire Effects Information System Plants
More info for the terms: fuel, fuel loading, litter

The half-life of Bishop pine litter is approximately 7.5 years. Biomass
and productivity varies between stands [43].

Fire reduces fuel loading and improves seedbed conditions for bishop
pine [27].
license
cc-publicdomain
bibliographic citation
Cope, Amy B. 1993. Pinus muricata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Key Plant Community Associations

provided by Fire Effects Information System Plants
More info for the terms: grassland, mesic, natural, tree

Bishop pine is frequently dominant in closed-cone pine forests
[15,27,37,42,44]. Stands are open with little or no understory on dry,
rocky sites, with a more dense understory on moist sites [15]. Bishop
pine also occurs in mesic border areas of woodlands and savannas [27].

In the northern part of its range, Bishop pine occurs in pure stands and
in redwood (Sequoia sempervirens), Douglas-fir (Pseudotsuga menziesii),
and pygmy forests [15,26,28,44]. In the southern portion of its
range, it is found in annual grassland, coastal sage scrub, and chaparral
communities. Scattered bisop pine stands often form a mosaic with these
communities [6,15,18,44].

Bishop pine is named as a dominant tree in the following published
classifications:

Terrestrial natural communities of California [15]
A vegetation classification system applied to southern California [36]
The southern California islands [37]
Vascular plant communities of California [42]
The closed-cone pines and cypress [44]

Associated canopy species not previously mentioned include Gowen cypress
(Hesperocyparis goveniana), Monterey cypress (H. macrocarpa), Tecate cypress
(H. forbesii), Mendocino cypress (H. pigmaea), Bolander pine (Pinus contorta
var. bolanderi), Monterey pine (Pinus radiata), and Pacific madrone
(Arbutus menziesii) [3,6,17,26,45]. Understory associates include glossyleaf
manzanita (Arctostaphylos nummularia), woollyleaf manzanita (Arctostaphylos
tomentosa), Pacific rhododendron (Rhododendron macrophyllum), California
huckleberry (Vaccinium ovatum), and salal (Gautheria shallon) [3,15,17,26,44].
license
cc-publicdomain
bibliographic citation
Cope, Amy B. 1993. Pinus muricata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Life Form

provided by Fire Effects Information System Plants
More info for the term: tree

Tree
license
cc-publicdomain
bibliographic citation
Cope, Amy B. 1993. Pinus muricata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Management considerations

provided by Fire Effects Information System Plants
More info for the term: genotype

When planting Bishop pine, the genotype of planted seeds or trees vs.
the genotype of existing populations should be compared in order to
preserve the genetic purity of Bishop pine. seeds of unknown origin
were sown in an area of northern California where two genotypically
distinct Bishop pine overlap in the mid-1960's. Genes from the unknown
source have been found in seeds from native trees. As a result, trees
from this site has lost their value for genetic and evolutionary studies
[31].
license
cc-publicdomain
bibliographic citation
Cope, Amy B. 1993. Pinus muricata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Phenology

provided by Fire Effects Information System Plants
More info on this topic.

Cones of Bishop pine open for pollination between April and June
[10,15,19]. Growth is initiated in the spring [15].
license
cc-publicdomain
bibliographic citation
Cope, Amy B. 1993. Pinus muricata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Post-fire Regeneration

provided by Fire Effects Information System Plants
More info for the terms: crown residual colonizer, root crown

Tree without adventitious-bud root crown
Crown residual colonizer (on-site, initial community)
license
cc-publicdomain
bibliographic citation
Cope, Amy B. 1993. Pinus muricata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Taxonomy

provided by Fire Effects Information System Plants
More info for the term: cone

The scientific name of Bishop pine is Pinus muricata D. Don [8,22,35].

Bishop pine rarely hybridizes with Monterey pine (P. radiata); timing of
cone opening usually differs in the two species [8,44].
license
cc-publicdomain
bibliographic citation
Cope, Amy B. 1993. Pinus muricata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Physical Description

provided by USDA PLANTS text
Tree, Evergreen, Monoecious, Habit erect, Trees without or rarely having knees, Tree with bark rough or scaly, Young shoots 3-dimensional, Buds resinous, Leaves needle-like, Leaves alternate, Needle-like leaf margins finely serrulate (use magnification or slide your finger along the leaf), Leaf apex acute, Leaves > 5 cm long, Leaves < 10 cm long, Leaves yellow-green above, Leaves yellow-green below, Leaves not blue-green, Leaves white-striped, Needle-like leaves triangular, Needle-like leaves twisted, Needle-like leaf habit erect, Needle-like leaves per fascicle mostly 2, Needle-like leaf sheath persistent, Twigs glabrous, Twigs viscid, Twigs not viscid, Twigs without peg-like projections or large fascicles after needles fall, Berry-like cones orange, Woody seed cones < 5 cm long, Woody seed cones > 5 cm long, Seed cones bearing a scarlike umbo, Umbo with obvious prickle, Bracts of seed cone included, Seeds brown, Seeds black, Seeds winged, Seeds unequally winged, Seed wings prominent, Seed wings equal to or broader than body.
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compiler
Stephen C. Meyers
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Aaron Liston
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Steffi Ickert-Bond
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Damon Little
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USDA PLANTS text

Pinus muricata

provided by wikipedia EN

Pinus muricata, the bishop pine, is a pine with a very restricted range: mostly in California, including several offshore Channel Islands, and a few locations in Baja California, Mexico. It is always on or near the coast.[2]

In San Luis Obispo County it is found alone or in stands scattered on the coastal mountains and hills from Morro Bay to Shell Beach. A few stands of the tree are seen on the hills above the Sycamore Canyon Resort in Avila Beach. Within the City of San Luis Obispo, the Terrace Hill Open Space has several scattered specimens. Bishop pine seems to prefer already disturbed, unvegetated areas where it probably faces less competition from oaks and shrubs.

The common name "bishop pine" resulted from the tree having been first identified near the Mission of San Luis Obispo in San Luis Obispo, California. This tree has a large number of common names and other prior scientific names, due primarily to numerous variant forms. Other English names that have occasionally been used are prickle cone pine, Obispo pine, Santa Cruz pine and dwarf marine pine.

Description

Pinus muricata is a coniferous evergreen tree growing to a height of 15 to 26 metres (49 to 85 ft),[3] rarely up to 34 metres (112 ft), with a trunk diameter of up to 1.2 metres (3.9 ft). The species is often smaller, stunted and twisted in coastal exposures. It is drought-tolerant and grows on dry, rocky soil.

The needles are in pairs, green to blue-green, and 8–16 cm (3–6.5 in) long. Cones occur in one to five clusters.[4] The cones are strongly reflexed down the branch, 5-to-10-centimetre-long (2.0 to 3.9 in) long; the scales are stiff, thin on the side of the cone facing the stem, but greatly thickened on the side facing away and with a stout 5 to 12 millimetres (0.20 to 0.47 in) spine; both features adaptive to minimise squirrel predation and fire damage to the cones. The cones remain unopened for many years until fire or strong heat causes them to open and release the seeds.[5]

Forms

There are two Pinus muricata forms:

  • a southern form with bright green needles
  • a northern form with dark blue-green needles.

The resin composition also differs. The dividing line between the two is very sharp, five miles (8 km) south of the boundary between Mendocino County and Sonoma County, California. Experimental attempts to hybridize the two forms have consistently failed, indicating that their taxonomic relationship may be more distant than the very small differences in appearance would suggest.

Ecology

Growing in Salt Point, California

Pinus muricata is found with several oak and cypress associates within the California Coast Ranges. There are also a number of common understory flora associates including sword fern, salal and western poison oak.[6] Notable occurrences of P. muricata is in association with Mendocino cypress as a pygmy forest on coastal terraces in Mendocino County and Sonoma County, including one location within Salt Point State Park. It is classified an endangered species in Mexico.

Uses

Pinus muricata has been used in plantations with resultant growth rates higher than in the wild, but with adverse impacts to biodiversity.

This plant has ornamental value, and is cultivated in parks and gardens. It has gained the Royal Horticultural Society's Award of Garden Merit.[7]

References

  1. ^ Farjon, A. (2013). "Pinus muricata". IUCN Red List of Threatened Species. 2013: e.T34058A2841776. doi:10.2305/IUCN.UK.2013-1.RLTS.T34058A2841776.en. Retrieved 11 November 2021.
  2. ^ "Pinus muricata" at the Encyclopedia of Life
  3. ^ Cope, Amy B. (1993). "Pinus muricata". Fire Effects Information System (FEIS). US Department of Agriculture (USDA), Forest Service (USFS), Rocky Mountain Research Station, Fire Sciences Laboratory.
  4. ^ W.L. Jepson, 1909
  5. ^ Moore, Gerry; Kershner, Bruce; Craig Tufts; Daniel Mathews; Gil Nelson; Spellenberg, Richard; Thieret, John W.; Terry Purinton; Block, Andrew (2008). National Wildlife Federation Field Guide to Trees of North America. New York: Sterling. p. 90. ISBN 978-1-4027-3875-3.
  6. ^ C.M. Hogan, 2008
  7. ^ RHS Plant Selector Pinus muricata AGM / RHS Gardening

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Wikipedia authors and editors
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wikipedia EN

Pinus muricata: Brief Summary

provided by wikipedia EN

Pinus muricata, the bishop pine, is a pine with a very restricted range: mostly in California, including several offshore Channel Islands, and a few locations in Baja California, Mexico. It is always on or near the coast.

In San Luis Obispo County it is found alone or in stands scattered on the coastal mountains and hills from Morro Bay to Shell Beach. A few stands of the tree are seen on the hills above the Sycamore Canyon Resort in Avila Beach. Within the City of San Luis Obispo, the Terrace Hill Open Space has several scattered specimens. Bishop pine seems to prefer already disturbed, unvegetated areas where it probably faces less competition from oaks and shrubs.

The common name "bishop pine" resulted from the tree having been first identified near the Mission of San Luis Obispo in San Luis Obispo, California. This tree has a large number of common names and other prior scientific names, due primarily to numerous variant forms. Other English names that have occasionally been used are prickle cone pine, Obispo pine, Santa Cruz pine and dwarf marine pine.

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Wikipedia authors and editors
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wikipedia EN