定义：The marine benthic biome (benthic meaning 'bottom') encompasses the seafloor and includes such areas as shores, littoral or intertidal areas, marine coral reefs, and the deep seabed.
定义：Hard framework, internal or external, which supports and protects softer parts of plant, animal or unicellular organism, and to which muscles usually attach in animals, includes skeletons (derived from Lawrence, 2005).
