^Just, W., A. Baumstark, A. Suss, A. Graphodatsky, W. Rens, N. Schafer, I. Bakloushinskaya et al. 2007. Ellobius lutescens: Sex determination and sex chromosome. Sexual Development 1:211-221.
^Arakawa, Y., C. Nishida-Umehara, Y. Matsuda, S. Sutou, and H. Suzuki. 2002. X-chromosomal localization of mammalian Y-linked genes in two XO species of the Ryukyu spiny rat. Cytogenetic and Genome Research 99:303-309.