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Lifespan, longevity, and ageing

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Maximum longevity: 9.8 years (captivity) Observations: One captive specimen lived 9.8 years (Richard Weigl 2005).
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AnAge articles

Без наслов ( англиски )

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Tamiasciurus and Sciurus likely diverged in the late Pliocene. Red squirrels are first reported in the Irvingtonian and are now known from more than 30 late Rancholabrean faunas from the central and eastern United States.

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Rubin, C. 2012. "Tamiasciurus hudsonicus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Tamiasciurus_hudsonicus.html
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Catherine Rubin, University of Alaska Fairbanks
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Link Olson, University of Alaska Fairbanks
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Catherine Kent, Special Projects
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Animal Diversity Web

Behavior ( англиски )

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Red squirrels have well-developed and extremely acute senses of smell, sight, and hearing. They are well known for their ability to communicate by calls. These calls consist of rattles, screeches, growls, buzzes and chirps and are often used in defense of territory or in response to threats from predators. During mating, red squirrels use low aggressive calls or territorial calls to drive off subordinate males. Red squirrels may even be able to recognize each other by individual call. Communication is important because of their territorality. They intensively use vocal communication to advertise these territories and to threaten other squirrels. Both vocal and scent marking aid in the recognition of individuals. Olfactory communication is important because it can leave long-lasting impressions which advertise if the territory is taken and males can discriminate between different olfactory signatures of other males. Another reason chemical communication is advantageous is because it may also reduce predation risk. Scent marking may also enable the squirrels to avoid unnecessary chases and fights by becoming known to their neighbors. Red squirrels may also be able to make predator-specific calls, but evidence of this is still very inconclusive. Red squirrels tend to produce a high-frequency call for aerial predators and a harsher, bark-type call for terrestrial predators. However, it is more commonly observed that these two call types are mixed.

Communication Channels: visual ; tactile ; acoustic ; chemical

Other Communication Modes: pheromones ; scent marks

Perception Channels: visual ; tactile ; acoustic ; chemical

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Rubin, C. 2012. "Tamiasciurus hudsonicus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Tamiasciurus_hudsonicus.html
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Catherine Rubin, University of Alaska Fairbanks
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Link Olson, University of Alaska Fairbanks
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Catherine Kent, Special Projects
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Animal Diversity Web

Conservation Status ( англиски )

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As of 2008, red squirrels are classified as Least Concern on the ICUN Red List and by the United States government. They are widespread and common, have suitable habitat throughout their range, and face no major threats. One subspecies, Mt. Graham red squirrels (Tamiasciurus hudsonicus grahamensis), is endangered according to the ICUN Red List. This subspecies is only found in southeast Arizona and its population is about 150 individuals.

US Federal List: endangered

CITES: no special status

State of Michigan List: no special status

IUCN Red List of Threatened Species: least concern

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Rubin, C. 2012. "Tamiasciurus hudsonicus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Tamiasciurus_hudsonicus.html
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Catherine Rubin, University of Alaska Fairbanks
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Link Olson, University of Alaska Fairbanks
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Catherine Kent, Special Projects
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Benefits ( англиски )

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Red squirrels cause considerable economic losses. They interfere with conifer reforestation by eating 60 to 100% of cone crops and directly damaging trees through bud consumption and bark stripping. Squirrels also cause damage to human property by nesting in homes and gnawing on household items. They may bite humans if provoked.

Negative Impacts: injures humans (bites or stings); household pest

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Rubin, C. 2012. "Tamiasciurus hudsonicus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Tamiasciurus_hudsonicus.html
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Catherine Rubin, University of Alaska Fairbanks
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Link Olson, University of Alaska Fairbanks
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Catherine Kent, Special Projects
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Animal Diversity Web

Benefits ( англиски )

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Red squirrels are the third-most commonly harvested furbearer in Canada, bringing in about $1 million annually. Each year in Minnesota, thousands of red squirrels are harvested for consumption by humans. Red squirrels are an important prey item for other economically important species like lynx (Lynx) and martens (Martes).

Positive Impacts: food ; body parts are source of valuable material

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Rubin, C. 2012. "Tamiasciurus hudsonicus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Tamiasciurus_hudsonicus.html
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Catherine Rubin, University of Alaska Fairbanks
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Link Olson, University of Alaska Fairbanks
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Catherine Kent, Special Projects
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Animal Diversity Web

Associations ( англиски )

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Red squirrels impact the forest ecosystem by dispersing seeds and fungi through caching and forgetting about or otherwise failing to return to food caches. The diversity and abundance of beneficial ectomycorrhizal fungi in these caches helps young trees acquire nutrients and grow. They limit the regrowth of trees by eating the seeds and inner tissues of the trees, which can cause significant damage to tree survival and value. Red squirrels provide a feeding opportunity for porcupines (Erethizontidae) during the winter by peeling away the bark of lodgepole pines. Their feeding habits also cause conifers to grow multiple tops, which reduces their timber value but increases suitable nest sites for many arboreal rodents (Rodentia) and passerine birds (Passeriformes). The increased availability of nest sites sustains species richness.

Red squirrels are host to a variety of endoparasites and ectoparasites. Endoparasitic species include 9 species of nematodes, 9 species of tapeworms including the genus (Hymenolepis). Other endoparasites include tularemia bacteria (Francisella tularensis) and Emmonsia crescens), and some kinds of protists like sarocysts (Sarocystis), and (Haplosporanigium). They are also vulnerable to fungal lung disease infection via adiaspiromycosis (Emmonsia parva). Ectoparasites of red squirrels include 31 species of mites, ticks, and chiggers (Glycyphagidae and Acarina), 25 species of fleas including Siphonaptera, Opisodasys robustus, Orchopeas caedens, Orchopeas neotomae, Orchopeas leucopus, Oropsylla idahoensis, Ceratophyllus vison. They may also carry botfly larvae (Cuterebra emasculator). Viruses that infect red squirrels are silverwater virus, California encephalitis virus, and Powassan virus.

Ecosystem Impact: disperses seeds

Commensal/Parasitic Species:

  • sucking lice (Anoplura)
  • nematodes (Nematoda)
  • tapeworms (Cestoda)
  • cestodes (Hymenolepis)
  • fleas (Siphonaptera)
  • fleas (Opisodasys robustus)
  • fleas (Orchopeas caedens)
  • fleas (Orchopeas neotomae)
  • fleas (Orchopeas leucopus)
  • fleas (Oropsylla idahoensis)
  • fleas (Ceratophyllus vison)
  • ticks and chiggers (Acarina)
  • mites (Glycyphagidae
  • sarocysts (Sarocystis)
  • botflies (Cuterebra emasculator)
  • tularemia (Francisella tularensis)
  • adiaspiromycosis (Emmonsia parva)
  • adiaspiromycosis (Emmonsia crescens)
  • protists (Haplosporanigium)
  • silverwater virus
  • California encephalitis virus
  • Powassan virus
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Rubin, C. 2012. "Tamiasciurus hudsonicus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Tamiasciurus_hudsonicus.html
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Catherine Rubin, University of Alaska Fairbanks
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Link Olson, University of Alaska Fairbanks
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Catherine Kent, Special Projects
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Animal Diversity Web

Trophic Strategy ( англиски )

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Red squirrels are primarily granivorous, but they are also opportunistic omnivores in the absence of mast foods. Primary diet items vary with habitat and include the seeds of conifers and other tree types detailed below. They live in a resource pulse system, where foods (conifers like white spruce, Picea glauca) exhibit extreme annual variation. They consume a wide variety of mushrooms, including at least 45 species in the Cascade Mountains alone. Secondary food items include tree buds and flowers, fleshy fruits, tree sap, bark, insects, and other animal materials such as bird eggs or young snowshoe hares (Lepus americanus). During winter, spring, and early summer, bark stripping and tree girdling occurs commonly to access phloem and cambial tissues. Red squirrels are highly selective in their foraging behavior, harvesting cones from the tree species with the highest seed energy per cone first and systematically working their way through species of conifers by energy density per cone.

Red squirrels are primarily larder hoarders. In late summer through autumn, they harvest cones and store them in one or a few central middens. Middens are a central hoard that is easy to defend from competitors and provides a moist, cool environment that prevents cones from opening. Middens vary in size and number depending on habitat, food availability, and individual squirrel. However, they contain enough food to last one to two seasons and are often used by several generations of squirrels In the eastern United States and Canada, red squirrels frequently engage in scatter hoarding, which is a system involving many small hoards instead of a large midden. This accounts for 85% of all hoards and more than 50% of all cones stored in this area The downside to this type of storage is that middens only provides enough food for about 37 days. Red squirrels have a great sense of smell which they use when looking for middens during the winter. Some individuals store their food caches underground and are able to locate these seeds, even under 4 meters of snow. Whenever they do not recover a stockpile of food, the seeds are left to germinate.

Red squirrels pilfer food from other squirrels, but the extent varies greatly by population. Some squirrels the Yukon Territory of Canada almost never pilfer. Mt. Graham Red Squirrels, on the other hand, pilfer 97% of the time and in Vermont pilfering occurred 25% of the time. Age, boldness, and population density may play a role in pilfering.

Animal Foods: birds; mammals; eggs; insects

Plant Foods: wood, bark, or stems; seeds, grains, and nuts; fruit; flowers; sap or other plant fluids

Other Foods: fungus

Foraging Behavior: stores or caches food

Primary Diet: herbivore (Granivore )

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Rubin, C. 2012. "Tamiasciurus hudsonicus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Tamiasciurus_hudsonicus.html
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Catherine Rubin, University of Alaska Fairbanks
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Link Olson, University of Alaska Fairbanks
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Catherine Kent, Special Projects
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Distribution ( англиски )

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Biogeographic Regions: nearctic (Native )

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Rubin, C. 2012. "Tamiasciurus hudsonicus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Tamiasciurus_hudsonicus.html
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Catherine Rubin, University of Alaska Fairbanks
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Link Olson, University of Alaska Fairbanks
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Catherine Kent, Special Projects
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Animal Diversity Web

Habitat ( англиски )

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Red squirrels occupy northern boreal coniferous forests abundant with conifer seeds, fungi, and interlocking canopies. This limits them to mountain ranges on the southern and eastern boundaries of their range. In the Rocky Mountains they have been found at elevations up to 2,500 ft (762 m). Populations of red squirrels occur in different habit conditions due to the vastness of their range. They occur in both temperate and polar environments and are considered to be primarily arboreal. They can be found in a mixed variety of forests including coniferous, deciduous, and mixed forests and they are also able to thrive in suburban and urban settings, as long as cool, coniferous forests with dense, interlocking canopies and abundant fungal resources are present.

Range elevation: 0 to 762 m.

Habitat Regions: temperate ; terrestrial

Terrestrial Biomes: taiga ; forest ; mountains

Other Habitat Features: urban ; suburban ; riparian

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Rubin, C. 2012. "Tamiasciurus hudsonicus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Tamiasciurus_hudsonicus.html
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Catherine Rubin, University of Alaska Fairbanks
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Link Olson, University of Alaska Fairbanks
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Catherine Kent, Special Projects
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Animal Diversity Web

Life Expectancy ( англиски )

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Red squirrels exhibit a type 3 survivorship curve in wild populations. This means that most of the mortality is associated with deaths of the young and only 25% survive longer than 1 year. Despite their small size, red squirrels are considered relatively long lived, with the oldest known squirrel in the wild reaching 10 years of age. The longest recorded lifespan in captivity is 9 years. Mature squirrels are 2 to 4 years old and older squirrels are considered to be over 5 to 6 years old. Their average lifespan in the wild is 5 years.

Range lifespan
Status: wild:
10 (high) years.

Range lifespan
Status: captivity:
9 (high) years.

Average lifespan
Status: wild:
5 years.

Average lifespan
Status: captivity:
9.8 years.

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Rubin, C. 2012. "Tamiasciurus hudsonicus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Tamiasciurus_hudsonicus.html
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Catherine Rubin, University of Alaska Fairbanks
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Link Olson, University of Alaska Fairbanks
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Catherine Kent, Special Projects
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Animal Diversity Web

Morphology ( англиски )

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Red squirrels differ from other tree squirrels by their deep reddish color, territorial behavior, and their smaller body size. They are less than 30% the size of grey squirrels (Sciurus carolinensis). While size and pelage color can vary geographically, they generally have a reddish back and white underside that is demarcated by dark lateral lines, which are especially visible in summer. A white eye ring is present year-round and tufted ears are during the winter. Variation in the dorsal surface color can range from reddish to ferruginous brown to olivaceous gray, usually with a distinctive reddish or brownish lateral band running down the back. The tail is smaller and flatter than that of other tree squirrels and varies in color from yellowish-gray to rusty red, with a band of black often extending the entire length of tail. Where its range borders that of Douglas squirrels (Tamiasciurus douglasii), red squirrels are distinguished by color of their pelage. The underside of red squirrels is all white or cream, whereas Douglas squirrels are rust colored or with a blackish wash. Tail hairs have yellowish to rusty tips with a black band in red squirrels, whereas those of Douglas squirrels are white-tipped with a black band, making them moderately easy to distinguish from a distance. Male and female red squirrels are very similar in appearance.

Some ecogeographic variation is believed to occur, so length and weight measurements may only hold true for the population(s) in which they were measured. Twenty-five subspecies of red squirrels are recognized. They differ mainly by range distribution. Mass tends to range from 197.3 to 282.2 g, with an average of 212.97. Their total length ranges from 270 to 385 mm, with an average of 327.5. Much of their length is their tail, which is 92 to 158 mm long. Red squirrels have an average hind foot length of 35 to 57 mm and the ear length is 19 to 31 m. Condylobasal length is 42 to 50 mm. Red squirrels have a basal metabolic rate of 166 cm^3 oxygen/hour.

Red squirrels generally experience two annual molts, although the tail molts only once per year. The spring molt occurs from late March through August and starts on the nose and front feet, and ends on the rump. The fall molt is from late August to early December and begins on the tail and then progresses to the rump and the head before ending on the legs and flank. Molt stage can be an important age indicator. Skulls can be aged by looking at dental characteristics such as visible wear and presence of permanent teeth. Their dental formula is I 1/1, C 0/0, P 1/1 or 2/1, M 3/3. The total number of teeth is 20 or 22 depending on the presence of upper P3.

Range mass: 197.3 to 282.2 g.

Average mass: 212.97 g.

Range length: 270 to 385 mm.

Average basal metabolic rate: 166 cm3.O2/g/hr.

Other Physical Features: endothermic ; homoiothermic; bilateral symmetry

Sexual Dimorphism: sexes alike

Average basal metabolic rate: 1.615 W.

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Rubin, C. 2012. "Tamiasciurus hudsonicus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Tamiasciurus_hudsonicus.html
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Catherine Rubin, University of Alaska Fairbanks
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Link Olson, University of Alaska Fairbanks
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Catherine Kent, Special Projects
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Animal Diversity Web

Associations ( англиски )

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Red squirrels are likely prey for a variety of animals including snakes, birds of prey, and carnivorous mammals. They are preyed upon by Cooper's hawks (Accipiter cooperii), northern goshawks (Accipiter gentilis), bald eagles (Haliaeetus leucocephalus), great gray owls (Strix nebulosa), great horned owls (Bubo virginianus), American kestrels (Falco sparverius), red-shouldered hawks (Buteo lineatus), northern harriers (Circus cyaneus), red-tailed hawks (Buteo jamaicensis), and sharp-shinned hawks (Accipiter striatus). Mammals that prey upon them them are American martens (Martes americana) and fishers (Martes pennanti), weasels (Mustela), mink (Neovison vison), as well as various canids like red foxes (Vulpes vulpes) and felids like lynx (Lynx canadensis). They are also preyed upon by timber rattlesnakes (Crotalus horridus). Humans hunt red squirrels for both their fur and meat.

Alarm calls are the most highly used anti-predator adaptations of red squirrels. Red squirrels tend to produce a high-frequency call for avian predators and a harsher bark-type call for terrestrial predators. However, it is more common that these two call types are mixed when any predator approaches. Red squirrels have a high survival rate even when subjected to heavy predation. It is very hard to catch and kill these animals because they are quite agile and take to the trees or thick vegetation to escape predators. Red squirrels are also fairly aggressive and when cornered, will not hesitate to defend themselves.

Known Predators:

  • Cooper's hawks (Accipiter cooperii)
  • northern goshawks (Accipiter gentilis)
  • bald eagles (Haliaeetus leucocephalus)
  • great gray owls (Strix nebulosa)
  • great horned owls (Bubo virginianus)
  • American kestrels (Falco sparverius)
  • red-shouldered hawks (Buteo lineatus)
  • northern harriers (Circus cyaneus)
  • red-tailed hawks (Buteo jamaicensis)
  • sharp-shinned hawks (Accipiter striatus).
  • American martens (Martes americana)
  • fishers (Martes pennanti)
  • weasels (Mustela)
  • mink (Neovison vison)
  • canids (Canidae)
  • red foxes (Vulpes vulpes)
  • felids (Felidae)
  • lynx (Lynx canadensis)
  • timber rattlesnakes (Crotalus horridus)
  • humans (Homo sapiens)
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Rubin, C. 2012. "Tamiasciurus hudsonicus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Tamiasciurus_hudsonicus.html
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Catherine Rubin, University of Alaska Fairbanks
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Link Olson, University of Alaska Fairbanks
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Catherine Kent, Special Projects
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Animal Diversity Web

Reproduction ( англиски )

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Red squirrels have a defined breeding season lasting 105 days that can occur either once or twice a year. They will mate in early spring from March to May and then again in August to early September. A second breeding period tends to occur in warmer areas of their range. Although mate pairings may occur, red squirrels are generally characterized as promiscuous. Animals in the best condition tend to breed more regularly and successfully than animals in poor condition.

Red squirrels exhibit a scramble competition mating system, in which the main costs to males are locating receptive females. Males typically invade the territory of females in estrus and pursue them in obvious mate chases. During mate chasing, a single dominant male actively pursues a female and drives off other subordinate males using calls or direct chase. Mounting and copulation usually lasts several minutes and occurs several times in the afternoon of the only day the female is receptive. Copulation is frequently initiated by the female and terminated by the male. Mating most often occurs on the ground or in the lower branches of trees. During mating, the male holds the female around the posterior abdomen while resting his head on her back. It is common for both males and females to engage in grooming of the genitals before and after copulation. The adults have no further contact after mating and the male returns to his territory.

Mating System: polygynandrous (promiscuous)

Red squirrels breed once or twice a year, depending on their geographic location. Populations in the south and east generally have two litters a year, one in spring and one in late summer. This pattern occurs all the way up to Quebec and has also occurred in British Colombia. In the northern extent of their range, breeding only occurs once a year, generally in spring. In the eastern United States, two breeding seasons per year is common, one in March and a second in late July. Harsher conditions at the northern latitudes and higher elevations of the squirrels' range most likely limit reproduction to a single breeding season. Interannual variation in female reproduction is also common; there was a halving in the number of breeding females over 2 years in British Columbia. Their reproductive cycle has adapted to the cyclical production of conifer cone output. Female estrus is highest in February to March and June to July. Pregnancy peaks in March to April and August to September in New York. In Colorado, conception generally occurs April to June.

Females are in estrus only 1 day per breeding season. Conception usually occurs within a few hours of mating. Gestation averages 35 days. Newborn young weigh 7.08 g on average. Litter sizes range from 1 to 8 offspring but averages 3.97. Altricial young are born without any hair except vibrissae and fine chin hairs, but develop quickly. The external auditory meatus is obvious at 18 days, eyes open at 26 to 35 days, and pelage is fully developed in only 40 days. Lactation occurs for the first 70 days. After that time, young are cast out to find their own territory. In some cases when the mother is in poor condition she will give part of her territory to her offspring. This increases the probability of survival for the offspring, increasing the overall fitness the mother. Young are active outside the nest in 7 weeks and fully independent shortly after weaning. Dentition is complete and external skeletal measurements reach adult size by 125 days.

Most red squirrel nests are constructed within 30 m of cone caches. Red squirrels prefer natural cavities, but due to lack of such resources in coniferous forests, they construct leaf nests or occasionally underground nests. The most important factors influencing nest-tree selection are tree diameter and branching structure, and the availability of canopy escape routes. Nests are found at heights of 2 to 20 m, and while nest material varies with habitat, they typically include grasses, mosses, inner cambium, shredded bark leaves, feathers, and fur.

Reproduction is dependent on resource abundance. Red squirrels live in a resource-pulse system, where the main food (in most cases the seeds of conifer species like white spruce) fluctuates annually, in some cases spanning three orders of magnitude between failure and mast years. Food available for reproduction in the spring is determined by the abundance of cones produced the previous year. The reliance on and the defense of individual caches allows females to have the potential to assess the level of stored food that is available for current reproduction. However, female squirrels do not have a smaller litter in years of crop failure, so reproduction is expected to be more costly during these years.

Age also affects reproductive cycles in both male and female red squirrels. Females and males are sexually mature at 1 year of age but are still developing. Despite the detriment to their own health by trying to breed and grow simultaneously, 1-year-old females tend to reproduce because they can achieve higher lifetime reproductive success than females delaying their first reproduction. This is also true for young males. Mature females tend to engage in a conservative reproductive strategy in order to allocate reproductive resources only when their own survival costs are maintained. This is not the case for females over 6 years old, who tend to sacrifice their own survival for reproduction because they are unlikely to be able to breed again the following year. Reproduction senescence occurs from 4 years of age onwards.

Breeding interval: Red squirrels breed once or twice a year, depending on their geographic location.

Breeding season: The time of breeding depends on their geographic location.

Range number of offspring: 1 to 8.

Average number of offspring: 3.97.

Average gestation period: 35 days.

Average weaning age: 70 days.

Average time to independence: 7 weeks.

Average age at sexual or reproductive maturity (female): 1 years.

Average age at sexual or reproductive maturity (male): 1 years.

Key Reproductive Features: seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; viviparous ; sperm-storing

Average birth mass: 7 g.

Average gestation period: 37 days.

Average number of offspring: 4.2.

Average age at sexual or reproductive maturity (female)
Sex: female:
342 days.

Females raise the young without any help from males. Gestation averages 35 days and lactation is 70 days. After this the young are no longer dependent on the mother and the offspring relocate to find territories of their own. Mothers sometimes bequeath territories, giving part or all of their territory to one or more of their offspring. This increases the offspring’s’ probability of overwinter survival.

Parental Investment: altricial ; female parental care ; pre-fertilization (Provisioning, Protecting: Female); pre-hatching/birth (Provisioning: Female, Protecting: Female); pre-weaning/fledging (Provisioning: Female, Protecting: Female); pre-independence (Provisioning: Female, Protecting: Female)

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cc-by-nc-sa-3.0
авторски права
The Regents of the University of Michigan and its licensors
библиографски навод
Rubin, C. 2012. "Tamiasciurus hudsonicus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Tamiasciurus_hudsonicus.html
автор
Catherine Rubin, University of Alaska Fairbanks
уредник
Link Olson, University of Alaska Fairbanks
уредник
Catherine Kent, Special Projects
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Animal Diversity Web

Associated Plant Communities ( англиски )

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More info for the terms: competition, forest, hardwood

The red squirrel inhabits coniferous forests, mixed conifer-hardwood and
occasionally hardwood forests, and rural woodlots [6,19,13].

In western North America red squirrels occur in white spruce (Picea
glauca) and black spruce (P. mariana) stands [13,34], Douglas-fir
(Pseudotsuga menziesii) forests [22] and pine (Pinus spp.) forests [41].

In Alberta red squirrels occur in quaking aspen (Populus
tremuloides)-balsam poplar (P. balsamea) stands with scattered white
spruce and black spruce [19]. They also occupy habitats dominated by
jack pine (Pinus banksiana) with occasional stands of white spruce.
Tamarack (Larix laricina) bogs and black spruce bogs are usually
occupied only for periods by juvenile red squirrels searching for
suitable territories [13,24].

In Wyoming red squirrels are present in continuous whitebark pine (P.
albicaulis), Engelmann spruce (Picea engelmannii)-subalpine fir (Abies
lasiocarpa), and lodgepole pine (Pinus contorta) forests [18].

In Colorado red squirrels favor lodgepole pine forests over open stands
of ponderosa pine (P. ponderosa) [13]. They are also found in
Douglas-fir and in stands transitional between Douglas-fir and ponderosa
pine. Their absence from ponderosa pine stands may be a result of
interspecific competition with Abert's squirrels (Sciurus aberti).
Outside the range of Abert's squirrel, red squirrels are more common in
ponderosa pine stands (i.e., in Black Hills National Forest, South
Dakota, and Medicine Bow National Forest, Wyoming) [11].

In Arizona and Utah red squirrels prefer fir and spruce forests and are
more rarely found in ponderosa pine forests [13]. They are closely
associated with old-growth Douglas-fir, white fir (A. concolor), and
blue spruce (Picea pungens) stands [27]. Red squirrels are common in
low-elevation mixed species forests in central Arizona [46].

In eastern North America red squirrels are abundant in mature hardwood
forests containing some mature spruce (Picea spp.), eastern hemlock
(Tsuga occidentalis), or balsam fir (A. balsamea), but are rare in pure
hardwood stands that do not contain oaks (Quercus spp.), hickories
(Carya spp.), or walnuts (Juglans spp.) [25,53]. In the mountains of
Virginia red squirrels occur in Table Mountain pine (Pinus pungens)-oak
forests [45].
лиценца
cc-publicdomain
библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Fire Effects Information System Animals

Common Names ( англиски )

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red squirrel
pine squirrel
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cc-publicdomain
библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Fire Effects Information System Animals

Conservation Status ( англиски )

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Information on state- and province-level protection status of animals in the
United States and Canada is available at NatureServe, although recent changes
in status may not be included.
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cc-publicdomain
библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Fire Effects Information System Animals

Cover Requirements ( англиски )

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More info for the terms: cover, natural, tree

Nesting: Red squirrels prefer natural cavities or abandoned woodpecker
holes for nesting. When these are not available a leaf or grass nest is
constructed near the top of a tree with dense foliage. Occasionally red
squirrels use abandoned burrows of other species [6]. Nest trees in
Arizona have crowns that interlock with two or more adjacent trees, and
average 14 inches (35.6 cm) d.b.h. [47]. In New York leaf nests are
built in deciduous trees and pines. Trees used for leaf nests ranged
from 4 to 16 inches (10.2-40.6 cm) in diameter, were 15 to 70 feet
(4.5-21.3 m) tall, and the nests were 12 to 60 feet (3.7-18.3 m) above
the ground. Cavity trees ranged from 12 to 36 inches (30.5-91.4 cm) in
diameter and were 20 to 60 feet (6-18.3 m) tall; entrance holes were 7 to
50 feet (2.1-15 m) above the ground [25].

Shelter: Nest cavities are sometimes used by one to several red
squirrels in severe weather [25]. In winter red squirrels tunnel in
deep snow to find food and escape severe weather [3,13].

Cone-bearing Trees and Food Cache Sites: Red squirrels in Arizona
require 9 to 25 large, mature, cone-bearing trees (or a larger number of
smaller cone-bearing trees) per territory [27,47]. Red squirrels use
moist, shaded areas for the central food cache. These areas are created
by groups of mature trees with some understory vegetation nearby. Large
trees in a group closely spaced in 0.1 acre (0.04 ha) or less are
favored. Canopy cover at 90 percent of cache sites is greater than 60
percent. One or more large snags, fallen logs, or live trees act as
support structures for the cache [47].

In Pennsylvania primary red squirrel feeding sites (not caches) were
associated with burrow entrances, overstory cone-bearing pines, and
other primary feeding sites. They tended to be away from nest trees [50].
лиценца
cc-publicdomain
библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Distribution ( англиски )

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The range of the red squirrel extends from Quebec and Ontario west to
Alaska; south in the Appalachian Mountains to Tennessee; and south in
the Rocky Mountains to New Mexico [6].
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cc-publicdomain
библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Food Habits ( англиски )

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More info for the terms: competition, cone, fresh, hardwood, seed, serotinous, tree

Red squirrels consume conifer seeds, nuts, buds, sap, tender leaves,
fruits, flowers, fungi, bird eggs, and small vertebrates [6]. In
captive trials white spruce seed alone sustained red squirrels for at
least 3 weeks. Red squirrels consumed seeds from an average of 144
cones per day per squirrel, probably more than they could consume in the
wild. Red squirrels lost weight on black spruce seed alone, and
preferred white spruce over black spruce seeds when presented with both
[2,3]. In Arizona each red squirrel was estimated to consume the seeds
of 42,000 to 131,000 pine cones per year [27,47]. On New Brunswick jack
pine sites, red squirrels consumed at least 47 cones per day [7].

In summer a larger array of foods is eaten, including elm (Ulmus spp.)
and maple (Acer spp.) seeds, raspberries (Rubus spp.) and other fruits,
and green cones of pines, Douglas-fir, hemlock, cedar, and larch (Larix
spp.). Some insects and nestling birds are also consumed [25]. In New
Hampshire red squirrels were observed preying on the eggs and young of
black-throated blue warblers (Dendroica caerulescens) [30]. Red
squirrels lap sap from trees damaged by yellow-bellied sapsuckers
(Sphyrapicus varius), and from other tree wounds [17]. They were also
observed licking sap and dried sugar from sap wells drilled by red-naped
sapsuckers (S. nuchalis) in Colorado [8]. In Vermont red squirrels were
observed biting holes in hardwood trees at periods associated with sap
flow. Sugar maple (Acer saccharum) was chosen more often than any other
tree. Rather than licking sap immediately, the red squirrels left the
fresh bite and returned at a later time, licking sap flows that had time
to evaporate into more concentrated sugar syrup or dried sugar. Sugar
spread by the researcher onto tree trunks did not immediately attract
red squirrels; it was inferred that red squirrels returned intentionally
to sap flows they had created [17].

The diet of the endangered Mount Graham red squirrel is not well known,
but is assumed to be similar to that of other red squirrels;
requirements include conifer seeds (mainly Engelmann spruce and
subalpine fir) and mushrooms [28].

In western North America cones are cached by red squirrels in a large
central location, often referred to as a midden (this type of food
storage is known as larderhoarding). Middens also contain other foods
such as seeds of other plant species, and mushrooms. Secondary middens
are also developed, usually at the base of a tree or tree cavity, but
are usually much smaller then the main midden [18,27]. Mushrooms are
sometimes left in tree branches and cached after they have dried [13].
It was suggested that larderhoarding may be a response to intense
predation in the winter; there is less movement involved in retrieving
cones from a cache than in searching for cones all winter [7].

In Wyoming red squirrels spent most of their foraging time in whitebark
pine forests finding and eating whitebark pine seed from cones or
recovering dropped whitebark pine seed. Cone harvesting may begin as
early as July 13 and cone caching usually begins by August 4. Red
squirrels often leave cones on the ground below the tree for up to 3
days before caching them [18]. In Colorado red squirrels appear to
prefer Douglas-fir cones over ponderosa pine cones; this may be a result
of interspecific competition with Abert's squirrels which require
ponderosa pine cones and bark [11]. In the Southwest commonly eaten and
preferred seeds include those of Douglas-fir, blue spruce, Engelmann
spruce, and white fir. Ponderosa pine seed is occasionally eaten in
Arizona [31].

In eastern North America red squirrels in conifer-dominated forests have
diets similar to those in western North America. In Maine red squirrels
remove northern white-cedar (Thuja occidentalis) branchlets and cones in
the fall, and consume branchlets with reproductive buds in the spring
[4]. Winter foods include terminal buds of conifers, boxelder (Acer
negundo) and red pine (Pinus resinosa) seeds, buds and sap of red maple
(A. rubrum), gray birch (Betula populifolia) and sugar maple, corn (Zea
mays), seeds of basswood (Tilia americana), sugar maple, silver maple
(A. saccharinum), Norway maple (A. platanoides), northern red oak
(Quercus rubra) and other oaks, hickory nuts, and butternuts (Juglans
cinera). Eastern hemlock (Tsuga canadensis) cones are a staple item in
some areas. Other winter foods include yellow-poplar (Liriodendron
tulipifera) and sycamore (Platanus occidentalis) seeds, fruits of
staghorn sumac (Rhus typhina) and smooth sumac (R. glabra), and bark.
In April and May red squirrels consume tree buds and flowers,
particularly those of American elm (Ulmus americana), sugar maple,
silver maple, Norway maple, northern red oak, and butternut [13,25,32].

In New Brunswick red squirrels are scatterhoarders rather than
larderhoarders; food is stored in small amounts in many places rather
than in one central midden. Red squirrels stored cones from both
serotinous and nonserotinous species: jack pine in central Ontario;
black, white, and red spruce (Picea rubens) in New Brunswick; and black
and red spruce in Maine. Cached cones are not sufficient for an entire
winter food supply; however, jack pine cones remain closed for at least
1 year and are retained on the tree for up to 7 years and therefore
provide a steady supply of winter food [7]. In Pennsylvania researchers
could not locate cone caches in marginal red squirrel habitat dominated
by aspens (Populus spp.). Red squirrels used a central location for
consuming cones; these feeding areas have been identified by the remains
of up to 103 cones [50].
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cc-publicdomain
библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Habitat-related Fire Effects ( англиски )

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More info for the terms: forest, low-severity fire, prescribed fire, presence, succession, tree

Kirkpatrick and Mosby [20] suggested that prescribed fire in southern
pine-hardwood sapling and older stages is unlikely to affect resident
tree squirrels. This may apply to red squirrels, since important
habitat includes mature trees unlikely to be adversely affected by
low-severity fire. Fire severe enough to cause basal fire wounds may
increase cavities available for food caches. Basal fire wounds are
unlikely to increase cavities useful as nests for red squirrels [20].

In Yellowstone National Park lodgepole pine stands monitored for
presence of birds and mammals during postfire succession, red squirrels
were only present in stands with closed canopies [42]. In north-central
Colorado red squirrels were not present on 8-year-old burned areas but
were present in adjacent unburned lodgepole pine stands [33].

The maintenance of many mature coniferous forest types is often
dependent on fire. Ponderosa pine, Douglas-fir, lodgepole pine,
whitebark pine, and spruces are either dependent on stand-replacing
fires for regeneration or on low-severity fires for maintenance. Even
though severe fire is immediately destructive of red squirrel habitat,
the long-term maintenance of most coniferous forests is dependent on
fire [57].
лиценца
cc-publicdomain
библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Habitat: Cover Types ( англиски )

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More info on this topic.

This species is known to occur in association with the following cover types (as classified by the Society of American Foresters):

1 Jack pine
5 Balsam fir
12 Black spruce
13 Black spruce-tamarack
14 Northern pin oak
15 Red pine
16 Aspen
18 Paper birch
19 Gray birch-red maple
20 White pine-northern red oak-red maple
21 Eastern white pine
22 White pine-hemlock
23 Eastern hemlock
24 Hemlock-yellow birch
25 Sugar maple-beech-yellow birch
26 Sugar maple-basswood
27 Sugar maple
30 Red spruce-yellow birch
31 Red spruce-sugar maple-beech
32 Red spruce
33 Red spruce-balsam fir
34 Red spruce-Fraser fir
35 Paper birch-red spruce-balsam fir
37 Northern white-cedar
38 Tamarack
40 Post oak-blackjack oak
51 White pine-chestnut oak
52 White oak-black oak-northern red oak
53 White oak
55 Northern red oak
78 Virginia pine-oak
107 White spruce
201 White spruce
202 White spruce-paper birch
204 Black spruce
205 Mountain hemlock
206 Engelmann spruce-subalpine fir
208 Whitebark pine
209 Bristlecone pine
210 Interior Douglas-fir
211 White fir
212 Western larch
213 Grand fir
215 Western white pine
216 Blue spruce
218 Lodgepole pine
219 Limber pine
223 Sitka spruce
224 Western hemlock
225 Western hemlock-Sitka spruce
227 Western redcedar-western hemlock
228 Western redcedar
230 Douglas-fir-western hemlock
237 Interior ponderosa pine
251 White spruce-aspen
253 Black spruce-white spruce
254 Black spruce-paper birch
лиценца
cc-publicdomain
библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Habitat: Ecosystem ( англиски )

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More info on this topic.

This species is known to occur in the following ecosystem types (as named by the U.S. Forest Service in their Forest and Range Ecosystem [FRES] Type classification):

FRES10 White-red-jack pine
FRES11 Spruce-fir
FRES14 Oak-pine
FRES15 Oak-hickory
FRES18 Maple-beech-birch
FRES19 Aspen-birch
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES22 Western white pine
FRES23 Fir-spruce
FRES24 Hemlock-Sitka spruce
FRES25 Larch
FRES26 Lodgepole pine
лиценца
cc-publicdomain
библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Habitat: Plant Associations ( англиски )

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More info on this topic.

This species is known to occur in association with the following plant community types (as classified by Küchler 1964):

More info for the terms: bog, forest

K001 Spruce-cedar-hemlock forest
K002 Cedar-hemlock-Douglas-fir forest
K004 Fir-hemlock forest
K005 Mixed conifer forest
K008 Lodgepole pine-subalpine forest
K011 Western ponderosa forest
K012 Douglas-fir forest
K013 Cedar-hemlock-pine forest
K014 Grand fir-Douglas-fir forest
K015 Western spruce-fir forest
K016 Eastern ponderosa forest
K017 Black Hills pine forest
K018 Pine-Douglas-fir forest
K019 Arizona pine forest
K020 Spruce-fir-Douglas-fir forest
K021 Southwestern spruce-fir forest
K022 Great Basin pine forest
K028 Mosaic of K002 and K026
K093 Great Lakes spruce-fir forest
K094 Conifer bog
K095 Great Lakes pine forest
K096 Northeastern spruce-fir forest
K097 Southeastern spruce-fir forest
K099 Maple-basswood forest
K100 Oak-hickory forest
K102 Beech-maple forest
K103 Mixed mesophytic forest
K104 Appalachian oak forest
K106 Northern hardwoods
K107 Northern hardwoods-fir forest
K108 Northern hardwoods-spruce forest
K111 Oak-hickory-pine forest
лиценца
cc-publicdomain
библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Habitat: Rangeland Cover Types ( англиски )

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More info on this topic.

This species is known to occur in association with the following Rangeland Cover Types (as classified by the Society for Range Management, SRM):

More info for the terms: forb, hardwood, shrubland

109 Ponderosa pine shrubland
409 Tall forb
809 Mixed hardwood and pine
лиценца
cc-publicdomain
библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Management Considerations ( англиски )

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More info for the terms: cone, cover, formation, natural, seed, selection, tree

Red squirrels are considered pests in some areas where they consume a
large proportion of available conifer seeds. It has been suggested that
they interfere with natural regeneration of conifers by removing most of
the seed crop; this is particularly detrimental in years with poor cone
crops [12,13,51]. Cached seed and cones do not usually germinate; a
seed fungus (Caloscypha fulgens), which appears to prevent germination
and rotting of seeds, is encouraged by the moist, cool conditions
usually found in red squirrel cache sites [53,54]. In Newfoundland red
squirrels are a recent introduction and have become a serious threat to
black spruce cone harvests for commercial seed production [48]. Red
squirrels clip (and often consume) terminal buds and peel the bark of
trees, occasionally girdling stems. Gnawing damage to trees sometimes
causes canker formation [13]. In Arizona red squirrels harvested more
than 80 percent of limber pine (Pinus flexilis) cones before the cones
opened (making the seeds unlikely to germinate even if they are not
consumed); in open sites where red squirrels were absent more than 70
percent of cones opened on the trees [9].

In Newfoundland harvesting black spruce cones in August or September is
recommended in order to anticipate removal of cones by red squirrels
[48]. In central British Columbia thinning of 20-year-old lodgepole
stands resulted in a reduction in red squirrel populations but did not
appreciably lower the amount of tree damage inflicted by red squirrels
gnawing on bark [41].

Red squirrels and other mycophagous species probably play an important
role in the dispersal of the spores of mycorrhiza-forming fungi and
therefore benefit conifer reproduction [26]. Red squirrel middens are a
source of conifer seed for forestry in some western states [12,13].

Habitat management: Red squirrels are dependent on mature conifers for
food and cover. Conifer plantings in suburban areas tend to encourage
red squirrels. Red squirrels will use nest boxes if they are placed in
suitable habitat [13]. Vahle and Patton [27,47] recommended maintenance
of areas with closely spaced groups of trees of different ages and sizes
for red squirrel habitat. Conifers larger than 15 inches (38 cm) d.b.h.
are necessary for cone production; at least three or four large trees
per acre are needed. Timber harvest using group selection promotes
habitats favored for red squirrel cache sites [47]. Reynolds and others
[31] listed specific management recommendations for red squirrels in the
Southwest in conjunction with maintenance of northern goshawk prey base.

Black bears (Ursus americanus) damaged trees adjacent to Mount Graham
red squirrel middens more often than at random locations. This damage
frequently kills the tree. Death of the tree results in loss of cone
production, but it also provides snags for nest cavities and storage
sites [39]. The endangered grizzly bear (U. arctos horribilis)
excavates red squirrel cone caches in whitebark pine stands; this is an
important grizzly bear food source in some areas. Grizzly bear habitat
management includes encouragement of both whitebark pine and red
squirrel [56].

Red squirrels are economically important furbearers in Canada [19,53].
More red squirrels are taken but the total value is less than that of
other furbearers [53].

Parasites and diseases of red squirrels were summarized by Layne [25]
and Flyger and Gates [13].
лиценца
cc-publicdomain
библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Occurrence in North America ( англиски )

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AK
AZ
CO
CT
DE
ID
IL
IN
IA
KY

ME
MD
MA
MI
MN
MT
NH
NJ
NM
NY

NC
ND
OH
OR
PA
RI
SC
SD
TN
UT

VT
VA
WA
WV
WI
WY

AB
BC
MB
NB
NF
NT
ON
PQ
SK
YT

лиценца
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библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Predators ( англиски )

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The main predators of red squirrels in Alberta are raptors, probably
principally northern goshawk (Accipiter gentilis) [24]. Other red
squirrel predators include red-tailed hawk (Buteo jamaicensis), red fox
(Vulpes vulpes), bald eagle (Halieaatus leucocephalus), marten (Martes
americana), lynx (Lynx lynx) and ermine (Mustela erminea) [13,44]. Lynx
prey slightly more heavily on red squirrels in summer than winter (2% of
lynx winter diet, 9% of summer diet) [13]. Red squirrels comprised 5.6
percent of 36 prey deliveries to northern goshawk nests and 17.5 percent
of pellets (in 8 nests). More than 100 red squirrels were taken per
season by a single pair of breeding northern goshawks [31].

In Yellowstone National Park martens use the subnivean zone to get to
their prey which includes red squirrel. Red squirrels comprised 25
percent by volume of marten scats [37].
лиценца
cc-publicdomain
библиографски навод
Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Preferred Habitat ( англиски )

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More info for the terms: cone, density, forest, seed

Red squirrels require mature coniferous trees as a source of cones and
seed [6]. The best cone production occurs in 200- to 300-year old
Douglas-fir, 40- to 300-year old white fir, and 150- to 200-year-old
Engelmann spruce. The best seed-producing stands of blue spruce (Picea
pungens) are 50 to 150 years old [31]. In central Colorado red
squirrels were present in closed stands of mature conifers [36]. In
west-central Colorado red squirrel caches were more abundant in
conifer-dominated stands than in mixed stands or in stands dominated by
quaking aspen [35]. In Ontario red squirrel tracks were more abundant
in uncut forest than in regenerating stands of various ages with brushy
understories [44].

In two study sites in central British Columbia red squirrels were twice
and over five times as abundant in unthinned stands of 20-year-old
lodgepole pine than in thinned stands. They were most abundant in
mature stands. It was suggested that the 20-year-old stands represented
marginal habitat that served as a sink area for excess juvenile red
squirrels [41].

Home Range: Red squirrel home range size varies with sex, age, and
habitat quality. Older red squirrels and females tend to have larger
home ranges. In lodgepole pine forests home ranges varied in size from
0.7 to 2.0 acres (0.3-0.8 ha); in mixed hardwood-conifer stands home
ranges varied from 6.5 to 11 acres (2.6-4.5 ha) [31]. Home ranges of
adult males in New York averaged 6.03 acres (2.4 ha) and ranged from
0.47 to 10.76 acres (0.2-4.3 ha) [25]. Davis [5] reported home ranges
of 3.2 acres (1.3 ha) for males and 3.7 acres (1.5 ha) for females.
Obbard [53] summarized other data on red squirrel home ranges.

Territoriality and Population Density: Red squirrels actively defend a
territory [18]. Gurnell [15] estimated that defended territories were
60 to 100 percent of the home range in lodgepole pine forests. In
British Columbia spruce-fir and hemlock-cedar (Thuja spp.) forests adult
territories averaged 2.2 acres (0.9 ha) at low elevations and 1.3 acres
(0.5 ha) at high elevations [52]. There are two types of defended
areas: most if not all red squirrels in Alberta defend a winter food
cache, which is usually abandoned during the summer, and some red
squirrels defend a "prime" territory which has an adequate food supply
year-round. One-quarter to one-third of adults defend a prime territory
[19]. In New Brunswick, where red squirrels are scatterhoarders instead
of larderhoarders, territories encompassing the entire food supply are
defended from conspecifics [7]. Smith [52] reported that defended red
squirrel territories in coniferous forests ranged from 0.5 to 3 acres
(0.2-1.2 ha).

Davis [5] reported red squirrel population density of 4 squirrels per 25
acres in mixed jack pine-black spruce stands and 23 red squirrels per 25
acres in white spruce forest. In southwestern Yukon Territory red
squirrel population densities of 27 per 25 acres (10 ha) and 17 per 25
acres were observed in white spruce forest interspersed with willow
(Salix spp.)-dominated meadows [29].

Many studies, reviewed by Klenner and Krebs [22], indicated that red
squirrel population density varies with cone crops. Zasada and others
[51] also suggested a relationship between red squirrel density and the
periodicity of white spruce cone production in Alaska since red
squirrels are dependent on white spruce cones for the majority of their
food. Rusch and Reeder [34] reported that summer populations fluctuated
between 67 and 151 red squirrels per 2,500 acres in mixed habitats.
Sullivan [40] reported that red squirrel population density in British
Columbia increased with food supplementation indicating that food
availability was limiting population density. In south-central British
Columbia food supplementation (feeders that were kept full of sunflower
[Helianthus spp.] seeds) in Douglas-fir and in white spruce forests
resulted in increases in red squirrel population densities. The amount
of change was greatest in the population occupying the Douglas-fir
stands, a low density population. Recruitment rates were very high
initially, but decreased with increasing population density. Each
population increased to approximately the same density, suggesting that
the factor regulating the maximum population density is not food supply
but territoriality. When feeders were removed, population density
declined to control levels in about 6 months [22].
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Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Regional Distribution in the Western United States ( англиски )

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More info on this topic.

This species can be found in the following regions of the western United States (according to the Bureau of Land Management classification of Physiographic Regions of the western United States):

5 Columbia Plateau
6 Upper Basin and Range
7 Lower Basin and Range
9 Middle Rocky Mountains
10 Wyoming Basin
11 Southern Rocky Mountains
12 Colorado Plateau
13 Rocky Mountain Piedmont
15 Black Hills Uplift
16 Upper Missouri Basin and Broken Lands
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Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Taxonomy ( англиски )

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The currently accepted scientific name for the red squirrel is
Tamiasciurus hudsonicus Erxleben. There are 25 accepted subspecies [13].
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Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Timing of Major Life History Events ( англиски )

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More info for the terms: density, litter, natural, tree

Diurnal Activity: Red squirrels are diurnal and active year-round. Peak
activity occurs 2 hours after sunrise and just before sunset, but
red squirrels are active most of the daylight hours [25].

Movement: Rusch and Reeder [34] reported that red squirrels move from
quaking aspen and pine stands to spruce stands in the spring. Fall
movements are largely those of juveniles moving away from denser
populations of adults in spruce stands.

Breeding Season: Red squirrel breeding season varies with latitude and
elevation. Generally, the breeding season peaks from March to August,
but breeding may begin as early as January and continue as late as
October. Mating is promiscuous [13,25].

Nesting and Litter Size: Female red squirrels prefer to nest in natural
tree cavities or abandoned woodpecker (Picidae) holes. If none are
available they build a leaf [25] or grass nest [27] in the crotch of a
limb high on a densely foliated tree, or use a fallen tree, rock pile,
underground burrow, or man-made structure. There are usually four or
five young per litter (range: one to eight). Generally there is only
one litter per year, but occasionally a female bears two litters in one
year [25].

Development of Young: Red squirrel young are born naked and blind. By
6 days there is noticeable fur growth and by 20 days there is a good
covering of dorsal fur. Their eyes open at 27 to 35 days. They are
mostly furred by about 8 weeks. The young red squirrels begin making
limited excursions from the nest at 6 to 7 weeks, and forage at a
greater distance by about 10 weeks [25]. Weaning occurs at about 7 to 8
weeks in New York [25] and about 10 weeks in Alberta [24]. Dispersal
usually begins at about 12 to 14 weeks [25]. Female red squirrels in
Canada usually do not breed until their second year. Males develop
scrotal testes their first spring but it is not known whether they breed
at that time [24].

Dispersal of Juveniles: Juvenile red squirrels disperse in the fall in
search of suitable territory [21]. In Alberta initial movements are
circular; unsuccessful juveniles usually return to the natal territory
until they are successful or are killed. Occasionally juveniles will
appropriate the mother's territory [24]. Survival of juveniles depends
on establishment of a territory with a food supply adequate for the
entire winter [22]. Juveniles surviving their first winter tended to be
relatively far away from the natal territory. They appeared to have
acquired larger territories than juveniles that died, with middens that
were constructed by previous owners [24].

Red squirrels tend to disperse and establish territories in relation to
major resources (mature cone-bearing trees) [22]. In south-central
British Columbia relocation of resident adult squirrels a substantial
distance from the study site allowed researchers to monitor red squirrel
recruitment rates. In the fall red squirrel population density
recovered to control levels on four out of six sites; immigrants were
mostly juveniles. In the spring, breeding red squirrel population
density recovered to control levels on five out of the six sites; these
sites were also colonized by juveniles in the fall [21].

Mortality and Longevity: In Alberta average annual mortality was 67
percent for juveniles, 34 percent for adult yearlings, and 61 percent
for older adults [19]. Larsen and Boutin [24] reported annual adult
survivorship for red squirrels in Alberta as 67 to 71 percent. Captive
red squirrels have lived at least 9 years and wild red squirrels have
lived at least 10 years [25].
лиценца
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Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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U.S. Federal Legal Status ( англиски )

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The Mount Graham red squirrel (T. h. ssp. grahamensis)is Endangered [45].
лиценца
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Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Use of Fire in Population Management ( англиски )

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More info for the term: fire regime

NO-ENTRY

FIRE REGIMES :
Find fire regime information for the plant communities in which this
species may occur by entering the species name in the FEIS home page under
"Find FIRE REGIMES".
лиценца
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Sullivan, Janet. 1995. Tamiasciurus hudsonicus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
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Gwiñver rous Norzhamerika ( бретонски )

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Gwiñver rous Norzhamerika (Tamiasciurus hudsonicus) a zo ur bronneg krigner hag a vev e Stadoù-Unanet Amerika ha Kanada.

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Esquirol vermell americà ( каталонски; валенсиски )

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L'esquirol vermell americà (Tamiasciurus hudsonicus) és una espècie de rosegador de la família dels esciúrids. Viu als Estats Units i gran part del Canadà. S'alimenta de llavors, pinyes, núcules, fruita i, de tant en tant, invertebrats i petits vertebrats. Els seus hàbitats naturals són els boscos de coníferes, mixtos o caducifolis, així com els arbustars i les zones amb vegetació secundària. Es creu que no hi ha cap amenaça significativa per a la supervivència d'aquesta espècie.[1]

Referències

 src= A Wikimedia Commons hi ha contingut multimèdia relatiu a: Esquirol vermell americà Modifica l'enllaç a Wikidata
  1. Linzey, A. V.; NatureServe (Hammerson, G.). Tamiasciurus hudsonicus. UICN 2008. Llista Vermella d'espècies amenaçades de la UICN, edició 2008, consultada el 19 octubre 2015.


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Esquirol vermell americà: Brief Summary ( каталонски; валенсиски )

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L'esquirol vermell americà (Tamiasciurus hudsonicus) és una espècie de rosegador de la família dels esciúrids. Viu als Estats Units i gran part del Canadà. S'alimenta de llavors, pinyes, núcules, fruita i, de tant en tant, invertebrats i petits vertebrats. Els seus hàbitats naturals són els boscos de coníferes, mixtos o caducifolis, així com els arbustars i les zones amb vegetació secundària. Es creu que no hi ha cap amenaça significativa per a la supervivència d'aquesta espècie.

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Gemeines Rothörnchen ( германски )

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Das Gemeine Rothörnchen (Tamiasciurus hudsonicus), auch Rothörnchen genannt, ist mit einer Körperlänge von 17 bis 23 und einer Schwanzlänge bis zu 16 Zentimetern ein eher kleiner Vertreter aus der Familie der Hörnchen.

Aussehen

Die Tiere haben eine rotbraune Fellfarbe am Rücken, der Kopf sowie der Bauch- und Brustbereich sind weiß gefärbt. Die Ohren sind nur mit einem sehr kurzen Fellbüschel versehen. Der Schwanz ist ebenfalls rotbraun und mit dünnen schwarzen und gelben Haaren durchzogen; die Spitze ist schwarz. Um die Augen ist ein auffälliger, weißer Ring vorhanden. Die sehr scharfen Krallen dienen den Tieren als Kletterhilfe. Das Gewicht der Einzeltiere beträgt zwischen 140 und 360 Gramm.

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Nahaufnahme des Kopfs des Rothörnchen

Verbreitung und Lebensraum

Das Gemeine Rothörnchen kommt in Alaska und Kanada mit Ausnahme des hohen Nordens vor. In den Vereinigten Staaten ist es in den Rocky Mountains, den Appalachen und südlich bis Pennsylvania beheimatet, dort bewohnt es die Nadelwälder.

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Verbreitungskarte des Rothörnchen

Lebensweise

Die tagaktiven Tiere sind Baumbewohner, sie fressen vor allem Samen, aber auch Nüsse, Eicheln, Zapfen, Knospen, Früchte, Pilze, gelegentlich auch Vogeleier, Nestlinge oder kleinere Mäuse. Für den Winter legt das Gemeine Rothörnchen bis zu einen Meter tiefe Vorratskammern am Boden an. Bei Gefahr geben die Tiere eine sehr schrillen Warnlaut, welcher an ein Zirpen erinnert, von sich. Die maximale Lebensspanne der Tiere beträgt ca. 10 Jahre.

 src=
Ein Rothörnchen im Winter
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Rothörnchen beim Verzehr eines Pinienzapfens

Fortpflanzung

Die erste Paarungszeit der Hörnchen ist im Februar oder März, die zweite im Juni oder Juli. Während der Paarungszeit streiten die Männchen sehr aggressiv um die Weibchen. Die Empfängniszeit des Weibchens beträgt nur einen Tag. Nach der Paarung verlassen die Männchen die Weibchen wieder. Nach einer Tragzeit von fünf Wochen kommen zwei bis sieben nackte und blinde Junge zur Welt, welche das Weibchen bis zu sieben Wochen säugt; die Augen öffnen die Jungen erst im Alter von zwei bis drei Wochen. Mit einem Alter von zwei Monaten verlassen die Jungen das elterliche Revier und suchen ein eigenes. Je nach Nahrungsverfügbarkeit kann das Weibchen im Jahr zwei Würfe bekommen. Die Geschlechtsreife erreichen die Tiere im Alter von einem Jahr.

Gefährdung

Da diese Art weit verbreitet ist und keine Bedrohungen bekannt sind, wird sie von der IUCN als ungefährdet (Least Concern) geführt. In Kanada werden die Tiere lokal als Lieferant für Pelze bejagt.

Belege

  • Sonnhild Bischoff, Dr. Helga Hofmann, Ursula Knopp, Angelika Lang: Neues Großes Tierlexikon in Farbe, Verlag: Compact Verlag München 2000, S. 324, ISBN 3-8174-5080-X.
  • Faszination Tier & Natur (Sammelordner): Karte: 296 Rothörnchen, Verlag: IMP/Meister Verlag GmbH München, Päckchen 86 056020 086 a.

Weblinks

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Gemeines Rothörnchen: Brief Summary ( германски )

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Das Gemeine Rothörnchen (Tamiasciurus hudsonicus), auch Rothörnchen genannt, ist mit einer Körperlänge von 17 bis 23 und einer Schwanzlänge bis zu 16 Zentimetern ein eher kleiner Vertreter aus der Familie der Hörnchen.

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Amerikaansk read iikhoarntsje ( западнофризиски )

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It Amerikaanske reade iikhoarntsje (Latynske namme: Tamiasciurus hudsonicus) is in sûchdier út it skift fan 'e kjifdieren (Rodentia), de famylje fan 'e iikhoarntsjes (Sciuridae) en it skaai fan 'e Noardamerikaanske reade iikhoarntsjes (Tamiasciurus), dat foarkomt yn in grut part fan Noard-Amearika. Dit bist wurdt pleatslik ek wol it dinne-iikhoarntsje (pine squirrel), de chickaree of de fairydiddle neamd. It Amerikaanske reade iikhoarntsje moat net betizze wurde mei it (gewoane) iikhoarntsje dat ek yn Fryslân foarkomt, en dat ek wol it (Jeropeeske of Jeraziatyske) reade iikhoarntsje neamd wurdt. De wittenskiplike soartenamme hudsonicus ferwiist nei de Hudsonbaai, dêr't it Amerikaanske reade iikhoarntsje yn 1771 foar it earst de soölooch Johann Christian Polycarp Erxleben ûnder eagen kaam.

Fersprieding

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It ferspriedingsgebiet fan it Amerikaanske reade iikhoarntsje.

It Amerikaanske reade iikhoarntsje komt foar yn in grut diel fan Noard-Amearika. Yn Kanada libbet er yn it grutste part fan it lân, útsein it Arktyske noarden en de eilannen Nijfûnlân, Kaap Bretoneilân, Prins Edwardeilân, de Magdalena-eilannen en Anticosti oan 'e eastkust en de Keninginne Sjarlotte-arsjipel oan 'e westkust. Teffens ûntbrekt er yn súdlik Alberta en yn 'e omkriten fan 'e stêd Vancouver, yn it súdwesten fan Britsk-Kolumbia. Yn 'e Feriene Steaten komt it Amerikaanske reade iikhoarntsje foar yn in grut diel fan 'e Rocky Mountains en de Grutte Dobbe, fan 'e Kanadeeske grins oant yn Arizona, mar hy ûntbrekt oan 'e westkust en op it grutste diel fan 'e Grutte Flakten. Wol komt er wer foar yn 'e krite fan 'e Grutte Marren, Nij-Ingelân, it legere noardeasten en yn 'e Appalachen oant yn Georgia.

 src=
In Amerikaansk read iikhoarntsje as túnornamint?

Amerikaanske reade iikhoarntsjes komme ek op ferskate plakken yn Jeropa foar as eksoat. Hjirby giet it wierskynlik om ûntsnapte eksimplaren út dieretunen of priveekolleksjes. Wierskynlik hat der him lykwols noch gjin fuortplantingspopulaasje fêstige. Yn Nederlân binne Amerikaanske reade iikhoarntsjes op ferskate plakken synjalearre, wêrûnder eastlik fan Arnhim, op it Waadeilân Teksel en op 'e Peel. Yn Fryslân hawwe der twa waarnimmings west yn Gaasterlân en ien by Dronryp. Yn 2018 wiene der fierders teminsten fjouwer Amerikaanske reade iikhoarntsjes yn 'e Ljouwerter Bosk, hoewol't der nei ferrin fan tiid noch mar ien waarnommen waard. De Rykstsjinst foar Undernimmend Nederlân (RVO), waard yn july fan dat jier foar de rjochter dage troch it Platfoarm Stopje Ynvasive Eksoaten, om 'e oerheid te twingen en fang de bisten út 'e Ljouwerter Bosk wei.[1] Op 21 augustus 2018 die de rjochter útspraak dat it lêste Amerikaanske reade iikhoarntsje, dat fan fans de namme 'Boefke' krigen hie, yn 'e Ljouwerter Bosk bliuwe mocht, om't er mar "beheinde aksjeradius" hie en him allinnich net fuortplantsje koe.[2]

Uterlike skaaimerken

It Amerikaanske reade iikhoarntsje hat trochinoar in kop-romplingte fan 16½-23 sm, mei in sturtlingte fan 9-16 sm en in gewicht fan 140-310 g. Hy is lytser as de oare beambewenjende iikhoarntsjes dy't yn Noard-Amearika foarkomme en hat in opfallende tekening, mei in readbrune rêch en in wite bealch, dy't faninoar skaat wurde troch in skerpe demarkaasjeline. De sturt hat krekt as by oare beambewenjende iikhoarntsjes de foarm fan in plomsturt, en wurdt û.m. brûkt om it lykwicht te hâlden by it klimmen en draven yn 'e beammen en as roer by it springen.

Biotoop

Amerikaanske reade iikhoarntsjes binne fanwegen harren dieet fan konifearesied sterk bûn oan it foarkommen fan nullebeammen. Sadwaande libje se fral yn nullewâlden en wurdt harren ferspriedingsgebiet mear beheind troch de noard- en súdgrins fan 'e taigasône as troch klimatologyske faktoaren. Inkeld oan 'e westkust fan 'e Feriene Steaten ûntbrekke se yn 'e nullewâlden, mei't dy dêr it domein binne fan it nau besibbe douglasiikhoarntsje (Tamiasciurus douglasii). Resint liket it Amerikaanske reade iikhoarntsje syn areaal ek út te wreidzjen nei mingde wâlden dy't foar it meastepart út leafbeammen besteane.

Hâlden en dragen

It Amerikaanske reade iikhoarntsje is inkeld oerdeis aktyf en hat in eksklusyf territoarium, dat er it hiele jier rûn fûleindich ferdigenet tsjin soartgenoaten, mei as iennichste útsûndering de peartiid. Hy makket syn nêst fan gers heech yn 'e beammen of soms yn holtes yn 'e stammen fan spjirren, popelieren of nutebeammen. Nêsten op 'e grûn binne tige seldsum. Elts eksimplaar ûnderhâldt ferskate nêsten yn syn territoarium, en wyfkes mei jongen ferhúzje dy geregeldwei fan it iene nei it oare nêst. Der binne in pear gefallen bekend fan Amerikaanske reade iikhoarntsjes dy't har nêst makken yn minsklike wenten en dêrby isolaasjemateriaal brûkten om it nêst te beklaaien.

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In Amerikaansk read iikhoarntsje ferkent syn om-en-by.

It wyfke fan it Amerikaanske reade iikhoarntsje is mar ien dei djoeisk, mar foarôfgeande dêroan bejout se har bûten har eigen territoarium, wat nei alle gedachten de mantsjes yn 'e omkriten fan har oansteande djoeiskens op 'e hichte stelt. Op 'e dei dat se djoeisk is, wurdt se neisitten troch ferskate mantsjes, dy't allegear besykje de konkurrinsje fuort te jeien. Dat slagget lykwols nea, en it wyfke pearet feitliks mei 4-16 ferskillende mantsjes, wat in frij opmerklik barren is yn 'e bistewrâld. (In mantsje dat mei mear as ien wyfke pearet komt withoefaak foar, mar in wyfke dat mei mear as ien mantsje pearet is folle seldsumer.) Nei in draachtiid fan 31-35 dagen smyt it wyfke dan in nêst fan 1-5 (mar meastal 3-4) jongen, dy't keal te wrâld komme, mei in gewicht fan sa'n 10 g. Dy ferlitte it nêst foar it earst as se 42 dagen of dêromtrint binne, wurde mei 70 dagen ôfwûn en binne mei 125 dagen folgroeid en selsstannich.

 src=
In swimmend Amerikaansk read iikhoarntsje.

De wyfkes kinne it jiers nei harren berte dielnimme oan 'e fuortplanting, mar guon wachtsje dêrmei oant se twa of trije jier binne. De measte wyfkes bringe mar ien nêst yn 't jier te wrâld, mar soms slane se in jier oer, wylst guon soms ek twa ferskillende nêsten jongen yn ien jier produsearje. Amerikaanske reade iikhoarntsjes hawwe trochinoar mar 22% kâns om it earste jier te oerlibjen. Dêrnei is de libbensferwachting lykwols trochinoar 2⅓ jier mei in maksimum fan 8 jier. Har wichtichste natuerlike fijannen binne de Kanadeeske lynks, reade lynks, prêrjewolf, Amerikaanske oehoe, (gewoane) hauk, readsturtmûzefalk, Amerikaanske krie, Amerikaanske marter, (gewoane) foks, grize foks, wolf en wezeling.

Fretten

Amerikaanske reade iikhoarntsjes frette benammentlik sieden fan nullebeammen, mar foegje op opportunistyske wize ek wol oare gaadlike saken oan har menu ta. Dêrûnder binne beamknoppen en -nullen, en fierders poddestuollen (wêrûnder guon dy't foar minsken deadlik giftich binne), skimmels en blêden fan 'e wylch, de popelier en de bearedrúf. Ek frette se inkeld wol fûgelaaien en sels wol jongen fan 'e Amerikaanske hazze. Fierwei it grutste part fan har fretten krije se lykwols út sieden dy't se út spjirre-apels helje. Dêrfan lizze se hjerstmis in foarrie oan op ien plak, dêr't se dan winterdeis op tarre. Om by it sied te kommen, moatte se de skobben fan 'e spjirre-apels ôfbrekke, en dy smite se dan op ien bulte, dy't in rúchskerne neamd wurdt, en dy't wol in meter yn trochsneed wurde kin.

 src=
In Amerikaansk read iikhoarntsje sil oan it miel.

Status

It Amerikaanske reade iikhoarntsje hat de IUCN-status fan "net bedrige", om't er yn syn ferspriedingsgebiet noch rûnom foarkomt en om't syn populaasje stabyl liket te wêzen.

 src=
In Amerikaansk read iikhoarntsje fan foarren sjoen.

Undersoarten

Der binne 24 erkende ûndersoarten fan it Amerikaanske reade iikhoarntsje:

  • Bailey's read iikhoarntsje (Tamiasciurus hudsonicus baileyi)
  • Dakotaansk read iikhoarntsje (Tamiasciurus hudsonicus dakotensis)
  • echt Amerikaansk read iikhoarntsje (Tamiasciurus hudsonicus hudsonicus)
  • Fremonts read iikhoarntsje (Tamiasciurus hudsonicus fremonti)
  • Kenai-iikhoarntsje (Tamiasciurus hudsonicus kenaiensis)
  • Minnesotaansk read iikhoarntsje (Tamiasciurus hudsonicus )
  • Mogolloniikhoarntsje (Tamiasciurus hudsonicus mogollonensis)
  • Richardsons read iikhoarntsje (Tamiasciurus hudsonicus richardsoni)
  • Saint Lawrence-iikhoarntsje (Tamiasciurus hudsonicus laurentianus)
  • súdlik read iikhoarntsje (Tamiasciurus hudsonicus dixiensis)
  • Tamiasciurus hudsonicus abieticola
  • Tamiasciurus hudsonicus gymnicus
  • Tamiasciurus hudsonicus grahamensis
  • Tamiasciurus hudsonicus lanuginosus
  • Tamiasciurus hudsonicus lychnuchus
  • Tamiasciurus hudsonicus loquax
  • Tamiasciurus hudsonicus pallescens
  • Tamiasciurus hudsonicus petulans
  • Tamiasciurus hudsonicus picatus
  • Tamiasciurus hudsonicus preblei
  • Tamiasciurus hudsonicus regalis
  • Tamiasciurus hudsonicus streatori
  • Tamiasciurus hudsonicus ventorum
  • Ungave-iikhoarntsje (Tamiasciurus hudsonicus ungavensis)

Keppelings om utens

Boarnen, noaten en referinsjes

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Amerikaansk read iikhoarntsje: Brief Summary ( западнофризиски )

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It Amerikaanske reade iikhoarntsje (Latynske namme: Tamiasciurus hudsonicus) is in sûchdier út it skift fan 'e kjifdieren (Rodentia), de famylje fan 'e iikhoarntsjes (Sciuridae) en it skaai fan 'e Noardamerikaanske reade iikhoarntsjes (Tamiasciurus), dat foarkomt yn in grut part fan Noard-Amearika. Dit bist wurdt pleatslik ek wol it dinne-iikhoarntsje (pine squirrel), de chickaree of de fairydiddle neamd. It Amerikaanske reade iikhoarntsje moat net betizze wurde mei it (gewoane) iikhoarntsje dat ek yn Fryslân foarkomt, en dat ek wol it (Jeropeeske of Jeraziatyske) reade iikhoarntsje neamd wurdt. De wittenskiplike soartenamme hudsonicus ferwiist nei de Hudsonbaai, dêr't it Amerikaanske reade iikhoarntsje yn 1771 foar it earst de soölooch Johann Christian Polycarp Erxleben ûnder eagen kaam.

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American red squirrel ( англиски )

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The American red squirrel (Tamiasciurus hudsonicus) is one of three species of tree squirrels currently classified in the genus Tamiasciurus, known as the pine squirrels (the others are the Douglas squirrel, T. douglasii, and the southwestern red squirrel, T. fremonti). The American red squirrel is variously known as the pine squirrel, North American red squirrel and chickaree. It is also referred to as Hudson's Bay squirrel, as in John James Audubon's work The Viviparous Quadrupeds of North America (hence the species name). The squirrel is a small, 200–250 g (7.1–8.8 oz), diurnal mammal that defends a year-round exclusive territory. It feeds primarily on the seeds of conifer cones, and is widely distributed across North America wherever conifers are common, except on the Pacific coast of the United States, where its cousin, the Douglas squirrel, is found instead.

The squirrel has been expanding its range into hardwood forests.[3]

Taxonomy

American red squirrels should not be confused with Eurasian red squirrels (Sciurus vulgaris); since the ranges of these species do not overlap, they are both commonly referred to as "red squirrels" in the areas where they are native. The specific epithet hudsonicus refers to Hudson Bay, Canada, where the species was first catalogued by Erxleben in 1771.[4] A recent phylogeny suggests the squirrels as a family can be divided into five major lineages. Red squirrels (Tamiasciurus) fall within the clade that includes flying squirrels and other tree squirrels (e.g., Sciurus).[5] There are 25 recognized subspecies of red squirrels.[6]

The southwestern red squirrel (T. fremonti) was long considered conspecific with T. hudsonicus, but a 2016 phylogenetic study found it to be a distinct species.[7]

Description

Red squirrels can be easily distinguished from other North American tree squirrels by their smaller size, 28–35 cm (11–14 in) total length (including tail), territorial behavior, and reddish fur with a white venter (underbelly).[6] Red squirrels are somewhat larger than chipmunks.

The Douglas squirrel is morphologically similar to the American red squirrels, but has a rust-colored venter and is restricted to the southwestern coast of British Columbia and in the Pacific Northwest of the United States. The ranges of the American red squirrel and the Douglas squirrel overlap in southern British Columbia, northwestern Washington and eastern Oregon.[8]

Distribution

Front view, Gatineau Park, Quebec

American red squirrels are widely distributed across the North American continent. Their range includes: most of Canada, excluding northern areas with no tree cover, the southern half of Alberta and the southwestern coast of British Columbia; the southern half of Alaska; the Rocky Mountains area of the United States, and northern half of the eastern United States.[6][9] American red squirrels are abundant and not of conservation concern throughout much of their range. American red squirrels were introduced on Newfoundland and have lived there in abundance for decades.[10] However, an isolated population of red squirrels in Arizona has experienced considerable declines in population size. In 1987, this portion of the population was listed as an endangered species.[11]

Fresh Red Squirrel tracks in snow.
Fresh Red Squirrel tracks in snow.

Behavior

Feeding

Eating nut, in Edmonton, Alberta
Video of a red squirrel eating, in Quebec
Large midden constructed by American red squirrels, Kenai National Wildlife Refuge, Alaska

American red squirrels are primarily granivores, but incorporate other food items into their diets opportunistically.[6] In Yukon, extensive behavioral observations suggest white spruce seeds (Picea glauca) comprise more than 50% of a red squirrel's diet, but squirrels have also been observed eating spruce buds and needles, mushrooms, willow (Salix sp.) leaves, poplar (Populus sp.) buds and catkins, bearberry (Arctostaphylos sp.) flowers and berries, and animal material such as bird eggs or even snowshoe hare leverets (young).[12] White spruce cones mature in late July and are harvested by red squirrels in August and September. These harvested cones are stored in a central cache and provide energy and nutrients for survival over the winter and reproduction the following spring. The fallen scales from consumed seed cones can collect in piles, called middens, up to twelve meters across.[13] White spruce exhibits two- to six-year masting cycles, where a year of superabundant cone production (mast year)[14] is followed by several years in which few cones are produced.[15] American red squirrel territories may contain one or several middens.

American red squirrels eat a variety of mushroom species, including some that are deadly to humans.[16]

Reproduction

American red squirrels are spontaneous ovulators.[17][18] Females enter estrus for only one day, but venture from their territory prior to ovulation, and these exploratory forays may serve to advertise their upcoming estrus. On the day of estrus, females are chased by several males in an extended mating chase. Males compete with one another for the opportunity to mate with the estrous female. Estrous females will mate with 4 to 16 males. Gestation has been reported to range from 31 to 35 days.[19] Females can breed for the first time at one year of age, but some females delay breeding until two years of age or older. Most females produce one litter per year, but in some years reproduction is skipped, while in other years some females breed twice. Litter sizes typically range from one to five, but most litters contain three or four offspring. Offspring are pink and hairless at birth and weigh about 10 g. Offspring grow at approximately 1.8 g per day while nursing, and reach adult body size at 125 days. They first emerge from their natal nests at around 42 days, but continue to nurse until approximately 70 days.

Nests are most commonly constructed of grass in the branches of trees. Nests are also excavated from witches' broom – abnormally dense vegetative growth resulting from a rust disease – or cavities in the trunks of spruce, poplar, and walnut trees. American red squirrels rarely nest below ground. Each individual squirrel has several nests within its territory, and females with young move them between nests. Some behavior has been reported within human dwellings using insulation as nest material.

A three-year study of a population of red squirrels in southwest Yukon reported female red squirrels showed high levels of multiple-male mating and would even mate with males with similar genetic relatedness. The relatedness of parents had no effect on the neonatal mass and growth rate of their offspring, nor did it affect the survival rate of offspring to one year of age.[20]

Sociality

Red squirrels are highly territorial and asocial with very few non-reproductive physical interactions (0.6% of all recorded behaviours in one 19-year study).[21] The majority of physical interactions are in male-female matings and between females and their offspring before the offspring disperse to their own territories (see Dispersal and survival). The non-reproductive physical interactions recorded were all instances of chasing an intruder from a territory.[21]

Dispersal and survival

Red squirrel swimming

If juvenile American red squirrels are to survive their first winter, they must acquire a territory and midden. They can acquire a territory by competing for a vacant territory, creating a new territory or by receiving all or part of a territory from their mothers. This somewhat rare (15% of litters) female behavior is referred to as breeding dispersal or bequeathal, and is a form of maternal investment in offspring.[22] The prevalence of this behavior is related to the abundance of food resources and the age of the mother. In some cases, females will acquire additional middens prior to reproduction, which they later bequeath to their offspring.[23] Offspring that do not receive a midden from their mother typically settle within 150 m (3 territory diameters) of their natal territory.[22] Observations suggest that male red squirrels have environmentally induced, alternative reproductive strategies that result in higher incidences of sexually selected infanticide in years when food is plentiful.[24]

American red squirrels experience severe early mortality (on average only 22% survive to one year of age). The survival probability, however, increases to age three, when it begins to decrease again. Females that survive to one year of age have a life expectancy of 3.5 years and a maximum lifespan of 8 years.[25] Increased maternal attention is correlated with increased offspring growth rate and higher lifetime reproductive success.[26]

Chief predators include Canada lynx (Lynx canadensis), bobcat (Lynx rufus), coyote (Canis latrans), great horned owl (Bubo virginianus), northern goshawk (Accipiter gentilis), red-tailed hawk (Buteo jamaicensis), American crow (Corvus brachyrynchos), American marten (Martes americana), pacific marten (Martes caurina), red fox (Vulpes vulpes), gray fox (Urocyon cinereoargenteus), wolf (Canis lupus), and weasel (Mustela sp.).[27]

References

  1. ^ Cassola, F. (2017) [errata version of 2016 assessment]. "Tamiasciurus hudsonicus". IUCN Red List of Threatened Species. 2016: e.T42587A115192299. doi:10.2305/IUCN.UK.2016-3.RLTS.T42587A22250817.en. Retrieved 19 February 2022.
  2. ^ Thorington, R.W. Jr.; Hoffmann, R.S. (2005). "Family Sciuridae". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: a taxonomic and geographic reference (3rd ed.). The Johns Hopkins University Press. pp. 754–818. ISBN 978-0-8018-8221-0. OCLC 26158608.
  3. ^ Goheen, Jacob R.; Swihart, Robert K.; Robins, James H. (2003). "The anatomy of a range expansion: Changes in cranial morphology and rates of energy extraction for North American red squirrels from different latitudes". Oikos. 102: 33–44. doi:10.1034/j.1600-0706.2003.12407.x.
  4. ^ Woods, S.E.J. (1980). The Squirrels of Canada. National Museums of Canada.
  5. ^ Mercer, J. M.; Roth, V. L. (2003). "The Effects of Cenozoic Global Change on Squirrel Phylogeny" (PDF). Science. 299 (5612): 1568–72. Bibcode:2003Sci...299.1568M. doi:10.1126/science.1079705. PMID 12595609. S2CID 40366357.
  6. ^ a b c d Steele, M.A. (1998). "Tamiasciurus hudsonicus" (PDF). Mammalian Species (586): 1–9. doi:10.2307/3504443. JSTOR 3504443. S2CID 253900287. Archived from the original (PDF) on 2016-03-04. Retrieved 2017-01-06.
  7. ^ Hope, Andrew G.; Malaney, Jason L.; Bell, Kayce C.; Salazar-Miralles, Fernando; Chavez, Andreas S.; Barber, Brian R.; Cook, Joseph A. (2016-07-01). "Revision of widespread red squirrels (genus: Tamiasciurus) highlights the complexity of speciation within North American forests". Molecular Phylogenetics and Evolution. 100: 170–182. doi:10.1016/j.ympev.2016.04.014. PMID 27083861.
  8. ^ Steele, Michael A. & Koprowski, John L. (2001). North American Tree Squirrels. Smithsonian Institution. p. 149. ISBN 9781588341006.{{cite book}}: CS1 maint: multiple names: authors list (link)
  9. ^ Osgood, W.H. (1900). "Results of a biological reconnaissance of the Yukon River region. General account of the region. Annotated list of mammals. Annotated list of birds". North American Fauna. 19: 1–100. doi:10.3996/nafa.19.0001.
  10. ^ Payne, Neil F. (1976). "Red Squirrel Introduction to Newfoundland". Canadian Field-Naturalist. 90: 60–64.
  11. ^ Arizona Game and Fish Department. (2003). Tamiasciurus hudsonicus grahamensis. Archived 2010-12-30 at the Wayback Machine Heritage Data Management System, Arizona Game and Fish Department, Phoenix, AZ.
  12. ^ Willson, Mary F.; De Santo, Toni L.; Sieving, Kathryn E. (July 2003). "Red squirrels and predation risk to bird nests in northern forests". Canadian Journal of Zoology. 81 (7): 1202–1208. doi:10.1139/z03-096.
  13. ^ Baheyeldin, Khalid (2020-09-05). "Red Squirrel Harvesting and Storing White Spruce Cones". YouTube. Archived from the original on 2021-11-07.
  14. ^ Kelly, D (1994). "The evolutionary ecology of mast seeding" (PDF). Trends in Ecology & Evolution. 9 (12): 465–70. doi:10.1016/0169-5347(94)90310-7. PMID 21236924. Archived from the original (PDF) on 2010-05-25. Retrieved 2017-01-06.
  15. ^ Nienstaedt, Hans; Zasada, John C. (1990). "Picea glauca". In Burns, Russell M.; Honkala, Barbara H. (eds.). Conifers. Silvics of North America. Washington, D.C.: United States Forest Service (USFS), United States Department of Agriculture (USDA). Vol. 1 – via Southern Research Station.
  16. ^ Wernert, Susan J. (1982). Reader's Digest North American wildlife (2. pr. ed.). Pleasantville, NY [u.a.]: Reader's Digest Assoc. ISBN 978-0-89577-102-5.
  17. ^ Layne, James N. (1954). "The biology of the red squirrel Tamiasciurus hudsonicus loquax in central New York". Ecological Monographs. 24 (3): 227–268. doi:10.2307/1948465. JSTOR 1948465.
  18. ^ Millar, John S. (1970). "The breeding season and reproductive cycle of the western red squirrel". Canadian Journal of Zoology. 48 (3): 471–473. doi:10.1139/z70-079.
  19. ^ Lair, H. (1985). "Length of gestation in the red squirrel, Tamiasciurus hudsonicus". Journal of Mammalogy. 66 (4): 809–810. doi:10.2307/1380818. JSTOR 1380818.
  20. ^ When It Comes to Female Red Squirrels, It Seems Any Male Will Do. Newswise (20 June 2008).
  21. ^ a b Gorrell, Jamieson C.; McAdam, Andrew G.; Coltman, David W.; Humphries, Murray M.; Boutin, Stan (2010). "Adopting kin enhances inclusive fitness in asocial red squirrels". Nature Communications. 1 (1): 22. Bibcode:2010NatCo...1...22G. doi:10.1038/ncomms1022. PMID 20975694. S2CID 205305747.
  22. ^ a b Berteaux, Dominique; Boutin, Stan (2000). "Breeding Dispersal in Female North American Red Squirrels" (PDF). Ecology. 81 (5): 1311. doi:10.1890/0012-9658(2000)081[1311:BDIFNA]2.0.CO;2. JSTOR 177210.
  23. ^ Boutin, S; Larsen, K. W.; Berteaux, D (2000). "Anticipatory parental care: Acquiring resources for offspring prior to conception". Proceedings of the Royal Society B: Biological Sciences. 267 (1457): 2081–2085. doi:10.1098/rspb.2000.1252. PMC 1690781. PMID 11416912.
  24. ^ Haines, Jessica A.; Coltman, David W.; Dantzer, Ben; Gorrell, Jamieson C.; Humphries, Murray M.; Lane, Jeffrey E.; McAdam, Andrew; Boutin, Stan (2018). "Sexually selected infanticide by male red squirrels in advance of a mast year" (PDF). Ecology. 99 (5): 1242–1244. doi:10.25316/ir-1436. PMID 29543323.
  25. ^ McAdam, Andrew G.; Boutin, Stan; Sykes, Ainsley K.; Humphries, Murray M. (2007). "Life histories of female red squirrels and their contributions to population growth and lifetime fitness". Écoscience. 14 (3): 362. doi:10.2980/1195-6860(2007)14[362:LHOFRS]2.0.CO;2. S2CID 86378466.
  26. ^ Taylor, Ryan W.; et al. (2020). "Attentive Red Squirrel Mothers Have Faster Growing Pups and Higher Lifetime Reproductive Success" (PDF). Behavioral Ecology and Sociobiology. 74 (6). doi:10.1007/s00265-020-02856-7. S2CID 219176132.
  27. ^ Rubin, Catherine. "Tamiasciurus hudsonicus (red squirrel)". Animal Diversity Web. Retrieved 2021-06-26.

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American red squirrel: Brief Summary ( англиски )

добавил wikipedia EN

The American red squirrel (Tamiasciurus hudsonicus) is one of three species of tree squirrels currently classified in the genus Tamiasciurus, known as the pine squirrels (the others are the Douglas squirrel, T. douglasii, and the southwestern red squirrel, T. fremonti). The American red squirrel is variously known as the pine squirrel, North American red squirrel and chickaree. It is also referred to as Hudson's Bay squirrel, as in John James Audubon's work The Viviparous Quadrupeds of North America (hence the species name). The squirrel is a small, 200–250 g (7.1–8.8 oz), diurnal mammal that defends a year-round exclusive territory. It feeds primarily on the seeds of conifer cones, and is widely distributed across North America wherever conifers are common, except on the Pacific coast of the United States, where its cousin, the Douglas squirrel, is found instead.

The squirrel has been expanding its range into hardwood forests.

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Tamiasciurus hudsonicus ( шпански; кастиљски )

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La ardilla roja americana (Tamiasciurus hudsonicus) es una especie de roedor de la familia de los Sciuridae. Vive en los Estados Unidos y gran parte de Canadá. Se alimenta de semillas, piñas, nueces, fruta y, de vez en cuando, de invertebrados y pequeños vertebrados. Sus hábitats naturales son los bosques de coníferas, mixtos o caducifolios, así como las zonas con vegetación secundaria. Se cree que no hay ninguna amenaza significativa para la supervivencia de esta especie.[3]

Referencias

  1. Cassola, F. (2016). Tamiasciurus hudsonicus. The IUCN Red List of Threatened Species doi 10.2305/IUCN.UK.2016-3.RLTS.T42587A22250817.en
  2. Thorington, R.W. Jr.; Hoffmann, R.S. (2005). «Family Sciuridae». En Wilson, D.E.; Reeder, D.M, eds. Mammal Species of the World: a taxonomic and geographic reference (3rd edición). The Johns Hopkins University Press. pp. 754-818. ISBN 0-8018-8221-4. OCLC 26158608.
  3. Linzey, A. V.; NatureServe (Hammerson, G.). «Tamiasciurus hudsonicus». Lista Roja de especies amenazadas de la UICN 2008 (en inglés). ISSN 2307-8235. Consultado el 2 de junio de 2018.

Galería de imágenes

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Tamiasciurus hudsonicus: Brief Summary ( шпански; кастиљски )

добавил wikipedia ES

La ardilla roja americana (Tamiasciurus hudsonicus) es una especie de roedor de la familia de los Sciuridae. Vive en los Estados Unidos y gran parte de Canadá. Se alimenta de semillas, piñas, nueces, fruta y, de vez en cuando, de invertebrados y pequeños vertebrados. Sus hábitats naturales son los bosques de coníferas, mixtos o caducifolios, así como las zonas con vegetación secundaria. Se cree que no hay ninguna amenaza significativa para la supervivencia de esta especie.​

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Tamiasciurus hudsonicus ( баскиски )

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Tamiasciurus hudsonicus Tamiasciurus generoko animalia da. Karraskarien barruko Sciurinae azpifamilia eta Sciuridae familian sailkatuta dago.

Erreferentziak

  1. (Ingelesez)Mammals - full taxonomy and Red List status Ugaztun guztien egoera 2008an
  2. Erxleben (1777) Sciuridae Syst. Regn. Anim. 416. or..

Ikus, gainera

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Tamiasciurus hudsonicus: Brief Summary ( баскиски )

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Tamiasciurus hudsonicus Tamiasciurus generoko animalia da. Karraskarien barruko Sciurinae azpifamilia eta Sciuridae familian sailkatuta dago.

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Punaorava ( фински )

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Punaorava (Tamiasciurus hudsonicus) on Pohjois-Amerikan pohjoisosissa elävä pienikokoinen oravalaji.[2][4] Se on yksi laajimmalle levinneistä ja reviiritietoisimmista oravalajeista Pohjois-Amerikassa.[1][2]

Ulkonäkö ja koko

 src=
Havupuiden siemenet ovat punaoravan pääravintoa.

Punaoravalla on solakka ja jäntevä ruumis, voimakkaat takajalat ja vahvat kynnet, joiden ansiosta se on taitava kiipeilemään puissa. Se painaa tavallisesti 140–250 grammaa. Kokonaispituus kuonosta hännänpäähän on 28–35 senttimetriä, josta hännän osuus on 9,5–15 senttimetriä. Häntä ei ole yhtä tuuhea kuin muilla pohjoisamerikkalaisilla oravalajeilla.[2]

Punaoravan väritys vaihtelee suuresti paitsi yksilöittäin, myös elinympäristön ja vuodenajan mukaan. Turkki on selästä ruskehtava tai punaruskea ja vatsasta valkoinen tai vaaleankeltainen. Kesällä vatsa- ja selkäpuolta erottaa toisistaan molemmissa kyljissä kulkeva tumma juova. Suuria, mustia silmiä ympäröivät valkoiset renkaat ja myös hännässä on usein valkoinen reunus.[2]

Levinneisyys

Punaorava elää Pohjois-Amerikan nearktiselta alueella, missä se on eräs laajimmalle levinneistä oravalajeista.[2] Sen levinneisyysalue ulottuu Alaskasta koko Kanadan halki Koillis-Yhdysvaltoihin ja Appalakkien vuoristoa pitkin Tennesseehen asti. Sitä tavataan myös Kalliovuorilla New Mexicon korkeudelle asti.[2][4] Punaoravasta on sen levinneisyysalueella määritetty 24 eri alalajia[3].

Elinympäristö

Punaorava viihtyy niin havu-, lehti- kuin sekametsissäkin. Sitä tavataan myös metsäisillä esikaupunkialueilla.[2]

Käyttäytyminen

 src=
Punaorava osaa myös uida.

Punaorava on yksin elävä päiväeläin, joka ei nuku talviunta. Aktiivisimmillaan se on kaksi tuntia auringonnousun jälkeen ja juuri ennen auringonlaskua. Se pesii vanhassa tikankolossa, ontossa puussa tai jossakin muussa elinalueellaan sijaitsevassa onkalossa. Jos sopivaa koloa ei löydy, se rakentaa pesänsä korkealle oksanhaaraan ruohoista ja lehdistä.[2][4]

Punaoravan elinalueen koko vaihtelee yksilön sukupuolen, iän ja ympäristön mukaan; yleensä sen pinta-ala on 1–2,4 hehtaaria.[2][4] Jos ravinto alkaa käydä vähiin, punaorava etsii itselleen uuden elinalueen. Etsiessään uutta elinaluetta se voi ylittää myös vesistöjä, sillä se on taitava uimari.[2] Levinneisyysalueensa pohjoisosissa se kaivaa talvisin lumeen käytäväverkostoja.[2][4]

Punaoravalla on erinomainen haju-, näkö- ja kuuloaisti. Se ääntelee paljon ja rähisee äänekkäästi reviirilleen tunkeutuville lajitovereille. Se pakenee saalistajiaan tiheikköihin tai puihin ja puolustautuu uhattuna sisukkaasti.[2]

Lisääntyminen

 src=
Mount Grahamilla elävä tumma alalaji on vaarassa kuolla sukupuuttoon.

Punaorava on promiskuiteettinen laji – sekä urokset että naaraat parittelevat usean kumppanin kanssa.[4] Lauhkeassa ilmastossa sillä on kaksi lisääntymiskautta, joista ensimmäinen sijoittuu lopputalveen ja toinen keskikesään. Kylmässä ilmastossa on vain yksi lisääntymiskausi kevätalvella. Tiineys kestää 33–35 vuorokautta, minkä jälkeen naaras synnyttää pesäänsä tavallisimmin 4 tai 5 sokeaa ja karvatonta poikasta, jotka painavat keskimäärin 7 grammaa. Poikaset kasvavat nopeasti ja vieroittuvat emostaan 7–8 viikon kuluttua syntymästä. Ne jättävät pesän 12–14 viikon ikäisinä ja lähtevät emonsa reviiriltä omille teilleen syntymänsä jälkeisenä syksynä.[2][4]

Poikaskuolleisuus on punaoravalla hyvin korkea: vain neljäosa poikasista selviää yhden vuoden ikäiseksi, jolloin ne saavuttavat täysikasvuisuuden ja sukukypsyyden.[2] Elinikä on luonnossa enimmillään 10 vuotta.[4] Punaoravan tavallisimpia saalistajia ovat pöllöt, haukat, kärpät, käärmeet, ketut, punailvekset, kanadanilvekset ja amerikannäädät.[2]

Ravinto

Punaorava on lähes kaikkiruokainen. Se käyttää ravinnokseen enimmäkseen havupuiden siemeniä, mutta myös muita siemeniä, hedelmiä, pähkinöitä, kaarnaa, silmuja, hirvieläinten pudottamia jättösarvia, matelijoita, hyönteisiä, mahlaa, männynkäpyjä, sieniä (myös ihmiselle myrkyllisiä sieniä), munia, linnunpoikasia, hiiriä ja jäniksenpoikasia.[2]

Punaorava varastoi siemeniä ja pähkinöitä talven varalle maan, lehtikasojen ja kivien alle. Tarkan hajuaistinsa avulla se pystyy paikallistamaan ruokakätkön 30 senttimetrin syvyydestä maan alta ja 4 metrin syvyydestä lumen alta. Siemeniä jää kuitenkin paljon myös maahan, minkä vuoksi punaoravalla on suuri merkitys puiden leviämiselle.[2]

Suojelu ja uhat

Kanadassa metsästetään joka vuosi 1–3 miljoonaa punaoravaa, joiden turkisten arvo on yhteensä noin miljoona dollaria. Tästä huolimatta laji on runsaslukuinen ja elinvoimainen lähes koko elinalueellaan. Kaakkois-Arizonassa sijaitsevalla Mount Grahamilla elävä alalaji T. h. grahamensis on kuitenkin äärimmäisen uhanalainen, ja sen kanta on enää noin 150 yksilöä. Sen elinympäristöä uhkaavat jatkuvat metsänhakkuut ja vuorelle rakennettu observatorio ja leirintäalue.[2]

Punaoravasta on sekä haittaa että hyötyä puiden lisääntymiselle. Se levittää puiden siemeniä ja niille hyödyllisiä sieni-itiöitä. Toisaalta se saattaa syödä havupuiden siemensadosta jopa kaksi kolmasosaa ja aiheuttaa huomattavia vahinkoja puuviljelmille ja siemenvarastoille. Joskus se voi myös kotiutua asuinrakennukseen ja jyrsiä kodin rakenteita tai aiheuttaa muuta tuhoa.[2]

Lähteet

  1. a b Linzey, A.V. & NatureServe (Hammerson, G.): Tamiasciurus hudsonicus IUCN Red List of Threatened Species. Version 2014.1. 2008. International Union for Conservation of Nature, IUCN, Iucnredlist.org. Viitattu 30.6.2014. (englanniksi)
  2. a b c d e f g h i j k l m n o p q r s t Tanya Dewey: Tamiasciurus hudsonicus – Red squirrel Animal Diversity Web. University of Michigan, Museum of Zoology. Viitattu 10.12.2011. (englanniksi)
  3. a b Wilson, Don E. & Reeder, DeeAnn M. (toim.): Tamiasciurus hudsonicus Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed). 2005. Bucknell University. Viitattu 7.2.2012. (englanniksi)
  4. a b c d e f g h Janet Sullivan: Tamiasciurus hudsonicus Fire Effects Information System. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory. Viitattu 10.12.2011. (englanniksi)
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Punaorava: Brief Summary ( фински )

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Punaorava (Tamiasciurus hudsonicus) on Pohjois-Amerikan pohjoisosissa elävä pienikokoinen oravalaji. Se on yksi laajimmalle levinneistä ja reviiritietoisimmista oravalajeista Pohjois-Amerikassa.

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Écureuil roux (Amérique du Nord) ( француски )

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Tamiasciurus hudsonicus

L'Écureuil roux américain (Tamiasciurus hudsonicus) est une espèce d'écureuils présente au Canada et aux États-Unis.

Description

Sa longueur est de 28 à 35 cm et il pèse entre 140 et 250g. On le remarque à sa couleur qui est blanc sur son ventre et son dos est plus ou moins roux avec une ligne noire séparant le tout. Il est aussi agile et nerveux.

Il ne faut pas le confondre avec l'écureuil roux d'Eurasie (Sciurus vulgaris).

Répartition

 src=
Répartition

L'écureuil roux d'Amérique du Nord est largement répandu à travers le continent nord américain. Son aire de répartition couvre une grande partie du Canada ainsi que plusieurs états américains.

Comportement

Relations sociales

Bien que cet écureuil vive en solitaire, une équipe de chercheurs canadiens (Kluane Red Squirrel Project) a constaté plusieurs cas d’adoptions de petits orphelins[1]. Étonnamment les orphelins adoptés avaient tous un lien de parenté assez proche avec leur mère adoptive, qui était leur tante, leur sœur ou leur grand-mère, ce qui implique que les écureuils roux connaissent les liens de parentés avec les autres écureuils qui les entourent.

Les chercheurs supposent que ces rongeurs, qui fréquentent peu leurs congénères, reconnaissent les individus à leurs cris et que, lorsqu’un cri familier ne se fait plus entendre, ils vont voir ce qu’il est advenu de la portée.

Alimentation et importance écologique

Comme son cousin européen Sciurus vulgaris, Tamiasciurus hudsonicus consomme en automne et hiver (et peut-être en été) des champignons, dont des champignons hypogés, c'est-à-dire dont la fructification est souterraine. Ce faisant, cette espèce contribue à en disséminer les spores et joue un rôle utile pour la forêt, car ce sont des champignons mycorhiziens essentiels pour des essences forestières majeurs. Elle contribue donc indirectement à la bonne mycorhization des arbres mais aussi à la dispersion des truffes et d'autres champignons.

Une étude[2] a porté sur le contenus du tractus digestif de 75 écureuils roux nord-américains et de 138 Grands Polatouches (Glaucomys sabrinus) capturés de novembre à février en forêt boréale au nord-ouest de l'Alberta. De grandes quantités spores de champignons épigés ont été observées, principalement des spores de Bolétales, Russulaceae et Cortinariaceae. Des spores d'ascomycètes hypogés (Elaphomyces et Tuber sp.) et de basidiomycètes (Gautiera, Hymenogaster, Hysterangium et Rhizopogon sp.) ont également été trouvées dans le tube digestif, en moindre quantité. Moins souvent on a trouvé des restes de sporocarpes coriaces ou carbonacés venant espèces d'Hymenochaetaless, Diatrypaceae, Xylariaceae et Bankeraceae, ainsi que quelques spores de rouilles (e.g. Phragmidium fusiforme) avec des conidies (e.g., Helicoma sp.) et des ascospores de microfungi ascomycètes (e.g. Sordariaceae). Par contre, aucun tissu lichénique n'a pu être trouvé, alors qu'une hypothèse admise était que ces écureuils pouvaient trouver avec les lichens abondants au sol et sur les arbres un complément alimentaire hivernal. Dans la même étude, 110 fumées fécales (excréments) d'écureuil volant du nord ont été collectées dans la même zone de juin à août. Leur analyse a montré que cet écureuil consomme aussi une importante quantité de champignons en été.

Reproduction

Liste des sous-espèces

 src=
Tamiasciurus hudsonicus grahamensis, une sous-espèce en voie de disparition.

Selon MSW :

  • T. h. hudsonicus
  • T. h. abieticola
  • T. h. baileyi
  • T. h. dakotensis
  • T. h. dixiensis
  • T. h. fremonti
  • T. h. grahamensis
  • T. h. gymnicus
  • T. h. kenaiensis
  • T. h. lanuginosus
  • T. h. laurentianus
  • T. h. loquax
  • T. h. lychnuchus
  • T. h. minnesota
  • T. h. mogollonensis
  • T. h. pallescens
  • T. h. petulans
  • T. h. picatus
  • T. h. preblei
  • T. h. regalis
  • T. h. richardsoni
  • T. h. streatori
  • T. h. ungavensis
  • T. h. ventorum

Notes et références

  1. (en) J. C. Gorrell, A. G. McAdam, D. W. Coltman, M. M. Humphries et S. Boutin, « Adopting kin enhances inclusive fitness in asocial red squirrels », Nat. Commun., vol. 1,‎ 2010 (ISSN , DOI )
  2. (en) R. S. Currah, E. A. Smreciu, T. Lehesvirta, M. Niemi, and K. W. Larsen ; Fungi in the winter diets of northern flying squirrels and red squirrels in the boreal mixedwood forest of northeastern Alberta ; Can. J. Bot. 78(12): 1514–1520 (2000) ; doi:10.1139/cjb-78-12-1514 ; NRC Canada

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Écureuil roux (Amérique du Nord): Brief Summary ( француски )

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Tamiasciurus hudsonicus

L'Écureuil roux américain (Tamiasciurus hudsonicus) est une espèce d'écureuils présente au Canada et aux États-Unis.

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Tamiasciurus hudsonicus ( италијански )

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 src=
Scoiattolo rosso che mangia una noce.

Lo scoiattolo rosso (Tamiasciurus hudsonicus Erxleben, 1777) è una specie di scoiattolo arboricolo del genere Tamiasciurus originaria del Nordamerica.

Tassonomia

Attualmente, gli studiosi riconoscono 24 sottospecie di scoiattolo rosso americano[3][4][5]:

Descrizione

Lo scoiattolo rosso è noto per il grido di richiamo che emette ogni qualvolta è disturbato: un acuto e prolungato ciurr-ciurr. La lunghezza della testa e del corpo insieme varia da 16,5 a 23 cm ed è dunque 2/3 delle dimensioni dello scoiattolo grigio orientale (Sciurus carolinensis)[3][4]. Il mantello varia dal bruno-fulvo al brunastro con una striscia rossastra che, partendo dalle orecchie, arriva fino alla punta della coda, e una linea nerastra lungo i fianchi; gli occhi sono cerchiati di bianco[3][4][6][7]. La coda è rivestita di peli gialli oppure bianchi, mentre le parti ventrali sono bianche[3][4][7]. D'inverno, il colore del mantello sbiadisce e la linea nera laterale, già assente negli individui giovani, sparisce, mentre le orecchie vengono protette da piccoli ciuffi di peli neri e rossastri[3][7]. Una specie molto simile, lo scoiattolo di Douglas (Tamiasciurus douglasii), si differenzia dallo scoiattolo rosso per la sfumatura color ruggine delle regioni ventrali[3][4][6][7].

Distribuzione e habitat

 src=
Scoiattolo rosso nel Québec.

Lo scoiattolo rosso occupa un areale molto vasto, che dall'Alaska e dal Québec si spinge a sud, attraverso le Montagne Rocciose, fino al Nuovo Messico e, attraverso gli Appalachi, fino alla Carolina del Sud[2][3][5][6][7]. È stato introdotto anche sull'isola di Terranova, presso la costa orientale del Canada[2]. La sottospecie T. h. grahamensis, isolata da tutte le altre, è ristretta al monte Graham, una vetta dei monti Pinaleño, nel sud-est dell'Arizona[2][4][5][6].

Gli scoiattoli rossi vivono essenzialmente nelle foreste decidue e sempreverdi, ma hanno una gran capacità di adattamento, per cui costruiscono la loro tana in qualsiasi riparo, attorno alle case, nelle siepi e nel sottosuolo[3]. Scavano gallerie anche sotto la neve e si servono persino di quelle abbandonate dalle marmotte e dai tamia[3]. Grazie alla loro capacità di adattamento a vivere sottoterra o sotto mucchi di pietre o di legname sono riusciti a sopravvivere al massiccio disboscamento delle foreste che ha ridotto l'ambiente vitale degli scoiattoli prettamente arboricoli[3].

I nidi vengono costruiti sulle biforcazioni dei rami, tra le foglie, nelle tane dei picchi, nelle cavità naturali dei vecchi alberi, nel sottosuolo, nel terreno asciutto o anche sotto pietre e ceppi[3]. Occasionalmente gli scoiattoli rossi si servono anche di vecchi nidi di corvi e di falchi come fondamenta per i loro nidi. Questi sono composti di tre strati: all'esterno da un intreccio di ramoscelli che portano ancora attaccate le foglie, all'interno da uno strato compatto di foglie pressoché impermeabile, mentre la camera interna, grande 10–15 cm, è rivestita di materiale più fine quale erba secca, muschio, penne e peli.

Gli scoiattoli rossi sono animali diurni, ma approfittano anche delle notti di luna per uscire all'aperto e sono laboriosi durante tutto l'inverno, rimanendo nei loro nidi solo durante il maltempo[3].

Biologia

Alimentazione

Gli scoiattoli rossi fanno un grande spreco di cibo, danneggiando e rifiutando più di quanto non ne mangino, e conservandone più di quanto ne possano consumare. La loro dieta è essenzialmente vegetariana con le pigne di pino che costituiscono la base della loro alimentazione durante tutto l'anno[3]. Essi le raccolgono dai rami in autunno, quando sono ancora verdi, e prima che possano maturare e cadere lontano[3]. In seguito le trasportano giù dall'albero e le immagazzinano a gruppi di 150, o anche più, sotto il suolo umido in modo che non possano maturare prima del necessario. Anche altri semi, quali nocciole e bacche, rientrano tuttavia nella loro dieta; a primavera, quando le gemme dell'olmo e dell'acero si staccano, gli scoiattoli rossi ne spaccano la corteccia per succhiarne il succo, così come spaccano le mele per mangiarne i semi, lasciando invece intatta la polpa[3]. Sono anche ghiotti di funghi e spesso li conservano fino a seccarli nelle spaccature degli alberi e nei ceppi[3]. Si nutrono anche della letale Amanita muscaria, dato che sembrano immuni al suo veleno[3]. Infine, ultime fra le loro preferenze sono la corteccia del pioppo comune e del pioppo tremulo, di cui si nutrono nei periodi di carestia, durante i quali non disdegnano altri alimenti vegetali e persino insetti, come ad esempio larve di scarafaggi snidate dal legno, carne putrefatta e, occasionalmente uova e uccellini e, talvolta, uccidono e mangiano persino i piccoli di alcuni conigli[3].

Riproduzione

Il corteggiamento inizia in primavera e talvolta fin dal mese di gennaio[3]. Al pari di altri scoiattoli, la caccia nuziale consiste in una gara in cui diversi scoiattoli rossi si inseguono su e giù per gli alberi e sul terreno, lanciando ognuno un richiamo sincopato, dolce e monotono. In seguito si uniscono e l'accoppiamento avviene in marzo o aprile a seconda delle condizioni ambientali[3]. Il maschio e la femmina collaborano alla costruzione del nido ma, completatolo, il primo non partecipa più alla vita familiare.

I piccoli, 4 o 5 in media per nidiata, partoriti dopo un periodo di gestazione di 40 giorni, sono ciechi e completamente privi di peluria, che spunta solo dopo una decina di giorni[3]. Vengono nutriti per cinque settimane e poi vengono cacciati via dalla madre che avrà una seconda nidiata in autunno.

Nemici

Linci rosse, visoni, falchi adulti e gufi catturano facilmente gli scoiattoli rossi, ma i loro veri nemici sono le martore americane che, in alcune zone, si nutrono quasi esclusivamente di scoiattoli rossi[3]. Infatti si arrampicano con la stessa abilità degli scoiattoli e li inseguono a balzi lungo i rami più sottili degli alberi.

La penuria di animali da pelliccia in molte zone ha provocato un incremento della caccia agli scoiattoli rossi, il cui manto viene utilizzato come guarnizione[3].

Conservazione

 src=
T. h. grahamensis.

Lo scoiattolo rosso è una specie ampiamente diffusa e comune che non sembra correre alcun rischio di estinzione[2]. In alcune aree, la specie può causare danni alle piantagioni di conifere, dal momento che ne strappa la corteccia e si nutre di gemme e semi, sebbene possa anche giocare un ruolo importante nel rimboschimento disperdendo e sotterrando i semi. In Canada, è una delle specie più cacciate per la pelliccia[3][6][7].

Tuttavia, sebbene nella sua interezza la specie non sia attualmente minacciata, la sottospecie del monte Graham è costituita solamente da una popolazione esigua, stimata nel 2009 a circa 250 esemplari[8], tutti relegati in un'unica area isolata[2][5][7]. Ritenuto estinto negli anni '50, lo scoiattolo rosso del monte Graham venne riscoperto negli anni '70[4], ma negli ultimi decenni il suo areale e la sua popolazione si sono ridotti sempre più[9]. La principale minaccia per questa sottospecie è costituita dalla perdita e dalla frammentazione dell'habitat dovuta alla raccolta del legname, allo sviluppo di reti stradali e di strutture turistiche, e dalla costruzione di un controverso complesso di osservatori[2][4][5][7][8], mentre siccità, incendi ed esplosioni di insetti possono solo peggiorare una situazione già compromessa[10][11]. La perdita dell'habitat potrebbe inoltre comportare la competizione con lo scoiattolo di Abert (Sciurus aberti), introdotto nella zona[5][10], e si teme che i mutamenti climatici possano ridurre ulteriormente l'habitat disponibile[10].

Note

  1. ^ (EN) D.E. Wilson e D.M. Reeder, Tamiasciurus hudsonicus, in Mammal Species of the World. A Taxonomic and Geographic Reference, 3ª ed., Johns Hopkins University Press, 2005, ISBN 0-8018-8221-4.
  2. ^ a b c d e f g (EN) Linzey, A.V. & NatureServe (Hammerson, G.) 2008, Tamiasciurus hudsonicus, su IUCN Red List of Threatened Species, Versione 2020.2, IUCN, 2020.
  3. ^ a b c d e f g h i j k l m n o p q r s t u v w x Steele, M.A. (1998) Tamiasciurus hudsonicus. Mammalian Species, 586: 1-9.
  4. ^ a b c d e f g h University of Arizona - Mount Graham Biology Program: Mount Graham Red Squirrel (Tamiasciurus hudsonicus grahamensis) (December, 2009).
  5. ^ a b c d e f Hafner, D.J., Yensen, E. and Kirkland Jr, G.L. (1998) North American Rodents: Status Survey and Conservation Action Plan Archiviato il 14 maggio 2012 in Internet Archive.. IUCN/SSC Rodent Specialist Group, IUCN, Gland, Switzerland and Cambridge, UK.
  6. ^ a b c d e Nowak, R.M. (1991) Walker's Mammals of the World. The Johns Hopkins University Press, Baltimore and London.
  7. ^ a b c d e f g h Feldhamer, G.A., Thompson, B.C. and Chapman, J.A. (2003) Wild Mammals of North America: Biology, Management, and Conservation. Second Edition. The Johns Hopkins University Press, Baltimore.
  8. ^ a b Arizona Game and Fish Department - Wildlife Blog: Wildlife News, November 9th, 2009 - Mount Graham red squirrel fall 2009 count announced Archiviato il 16 febbraio 2013 in Internet Archive. (December, 2009).
  9. ^ U.S. Fish and Wildlife Service. (1992) Mount Graham Red Squirrel Recovery Plan. U.S. Fish and Wildlife Service, Albuquerque, New Mexico.
  10. ^ a b c U.S. Fish and Wildlife Service. (2008) Mount Graham Red Squirrel (Tamiasciurus hudsonicus grahamensis). 5-Year Review: Summary and Evaluation. U.S. Fish and Wildlife Service, Phoenix, Arizona.
  11. ^ Koprowski, J.L., Alanen, M.I. and Lynch, A.M. (2005) Nowhere to run and nowhere to hide: response of endemic Mt. Graham red squirrels to catastrophic forest damage. Biological Conservation, 126: 491-498.

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Tamiasciurus hudsonicus: Brief Summary ( италијански )

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 src= Scoiattolo rosso che mangia una noce.

Lo scoiattolo rosso (Tamiasciurus hudsonicus Erxleben, 1777) è una specie di scoiattolo arboricolo del genere Tamiasciurus originaria del Nordamerica.

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Amerikaanse rode eekhoorn ( холандски; фламански )

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De Amerikaanse rode eekhoorn (Tamiasciurus hudsonicus) is een zoogdier uit de familie van de eekhoorns (Sciuridae). De wetenschappelijke naam van de soort werd voor het eerst geldig gepubliceerd door Erxleben in 1777.

Voorkomen

De soort komt voor in Canada en de Verenigde Staten. In 2015 werden enkele individuen ook in België waargenomen, waar ze van nature niet voorkomen.[2][3]

Bronnen, noten en/of referenties
  1. (en) Amerikaanse rode eekhoorn op de IUCN Red List of Threatened Species.
  2. Amerikaanse rode eekhoorn gevonden. Natuur.Flits. Natuurpunt (23 september 2015). Geraadpleegd op 23 september 2015.
  3. Amerikaanse Rode Eekhoorn - Tamiasciurus hudsonicus. Waarnemingen.be (23 september 2015). Geraadpleegd op 23 september 2015.
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Amerikaanse rode eekhoorn: Brief Summary ( холандски; фламански )

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De Amerikaanse rode eekhoorn (Tamiasciurus hudsonicus) is een zoogdier uit de familie van de eekhoorns (Sciuridae). De wetenschappelijke naam van de soort werd voor het eerst geldig gepubliceerd door Erxleben in 1777.

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Sosnowiórka czerwona ( полски )

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Sosnowiórka czerwona[3] (Tamiasciurus hudsonicus) – gatunek gryzonia z rodziny wiewiórkowatych[4] występujący w niemal całej Ameryce Północnej – od Alaski, przez Kanadę (aż do wybrzeży Atlantyku w Quebecu oraz Nowej Fundlandii i Labradorze) po Arizonę i Nowy Meksyk na południu oraz Georgię na południowym wschodzie Stanów Zjednoczonych[2].

Wygląd

Czerwono-brązowe futro z białym brzuchem.

Średnie wymiary
  • Długość ciała 17-23 cm,
  • Długość ogona 9-16 cm,
  • Masa ciała 140-360 g.

Tryb życia

Dzienny tryb życia, żyje samotnie w lasach iglastych. Żywi się roślinami, nasionami a także, jajami, małymi ptakami i ssakami.

Obejmuje 24 podgatunki. W Czerwonej księdze gatunków zagrożonych Międzynarodowej Unii Ochrony Przyrody i Jej Zasobów gatunek ten został zaliczony do kategorii LC (najmniejszej troski). Nie ma poważniejszych zagrożeń dla populacji tego gatunku[2].

Podgatunki

Wyróżnione podgatunki T. hudsonicus[4]

  • T. hudsonicus hudsonicus Erxleben, 1777
  • T. hudsonicus abieticola A. H. Howell, 1929
  • T. hudsonicus baileyi J. A. Allen, 1898
  • T. hudsonicus dacotensis J. A. Allen, 1894
  • T. hudsonicus dixiensis Hardy, 1942
  • T. hudsonicus fremonti Audubon i Bachman, 1853
  • T. hudsonicus grahamensis J. A. Allen, 1894
  • T. hudsonicus gymnicus Bangs, 1899
  • T. hudsonicus kenaiensis A. H. Howell, 1936
  • T. hudsonicus lanuginosus Bachman, 1839
  • T. hudsonicus laurentianus Anderson, 1942
  • T. hudsonicus loquax Bangs, 1896
  • T. hudsonicus lychnuchus Stone i Rehn, 1903
  • T. hudsonicus minnesota J. A. Allen, 1899
  • T. hudsonicus mogollonensis Mearns, 1890
  • T. hudsonicus palescens A. H. Howell, 1942
  • T. hudsonicus petulans Osgood, 1900
  • T. hudsonicus picatus Swarth, 1921
  • T. hudsonicus preblei A. H. Howell, 1936
  • T. hudsonicus regalis A. H. Howell, 1936
  • T. hudsonicus richardsoni Bachman, 1839
  • T. hudsonicus streatori J. A. Allen, 1898
  • T. hudsonicus ungavensis Anderson, 1942
  • T. hudsonicus ventorum J. A. Allen, 1898
Gilles Gonthier - Tamiasciurus hudsonicus (by) (1).jpg

Przypisy

  1. Tamiasciurus hudsonicus, w: Integrated Taxonomic Information System (ang.).
  2. a b c Tamiasciurus hudsonicus. Czerwona księga gatunków zagrożonych (IUCN Red List of Threatened Species) (ang.).
  3. Włodzimierz Cichocki, Agnieszka Ważna, Jan Cichocki, Ewa Rajska, Artur Jasiński, Wiesław Bogdanowicz: Polskie nazewnictwo ssaków świata. Warszawa: Muzeum i Instytut Zoologii Polskiej Akademii Nauk, 2015, s. 297. ISBN 978-83-88147-15-9.
  4. a b Wilson Don E. & Reeder DeeAnn M. (red.) Tamiasciurus hudsonicus. w: Mammal Species of the World. A Taxonomic and Geographic Reference (Wyd. 3.) [on-line]. Johns Hopkins University Press, 2005. (ang.) [dostęp 2015-11-28]
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Sosnowiórka czerwona: Brief Summary ( полски )

добавил wikipedia POL

Sosnowiórka czerwona (Tamiasciurus hudsonicus) – gatunek gryzonia z rodziny wiewiórkowatych występujący w niemal całej Ameryce Północnej – od Alaski, przez Kanadę (aż do wybrzeży Atlantyku w Quebecu oraz Nowej Fundlandii i Labradorze) po Arizonę i Nowy Meksyk na południu oraz Georgię na południowym wschodzie Stanów Zjednoczonych.

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Tamiasciurus hudsonicus ( португалски )

добавил wikipedia PT

Tamiasciurus hudsonicus, o esquilo-vermelho-americano, é uma das três espécies de esquilos de atualmente classificadas no gênero Tamiasciurus. São animais de médio porte (200 a 250 gramas), mamíferos diurnos que defendem um território exclusivo durante todo o ano. A dieta destes esquilos é especializada em sementes de coníferas. Como tal, eles estão amplamente distribuídos entre a América do Norte onde as coníferas são comuns, exceto na costa do Pacífico.[1]

Referências

  1. Goheen, J.R., R.K. Swihart, and J.H. Robins. 2003. The anatomy of a range expansion: changes in cranial morphology and rates of energy extraction for North American red squirrels from different latitudes. Oikos, 102: 33-44.
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Tamiasciurus hudsonicus: Brief Summary ( португалски )

добавил wikipedia PT

Tamiasciurus hudsonicus, o esquilo-vermelho-americano, é uma das três espécies de esquilos de atualmente classificadas no gênero Tamiasciurus. São animais de médio porte (200 a 250 gramas), mamíferos diurnos que defendem um território exclusivo durante todo o ano. A dieta destes esquilos é especializada em sementes de coníferas. Como tal, eles estão amplamente distribuídos entre a América do Norte onde as coníferas são comuns, exceto na costa do Pacífico.

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Amerikansk röd ekorre ( шведски )

добавил wikipedia SV


Amerikansk röd ekorre (Tamiasciurus hudsonicus[2][3][4][5][6][7]) är en däggdjursart som först beskrevs av Johann Christian Polycarp Erxleben 1777. Den ingår i släktet Tamiasciurus och familjen ekorrar.[8][9] IUCN kategoriserar arten globalt som livskraftig.[1]

Utseende

Arten liknar den europeiska ekorren (Sciurus vulgaris) i utseende men den är tydlig mindre och dessutom tillhör de olika släkten. Individer av amerikansk röd ekorre når en kroppslängd (huvud och bål) av 16,5 till 23 cm, en svanslängd av 9 till 16 cm och en vikt av 195 till 280 gram.[10] Pälsens färg på ryggen varierar mellan rödbrun och brun medan buken är vitaktig. Under sommaren är kanten mellan den bruna och den vita pälsen ofta svartaktig. Den gulgråa till rödbruna svansen är inte lika yvig som hos andra arter av släktet Tamiasciurus. Kring ögonen har arten en iögonfallande vit ring.[10]

Utbredning och habitat

Amerikansk röd ekorre förekommer i stora delar av Nordamerika från Alaska över Kanadas tempererade delar till Stora slätterna i USA och östra USA. I centrala USA sträcker sig utbredningsområdet söderut till Arizona och New Mexico. I östra USA ligger utbredningsområdets södra gräns i nordvästra Georgia och södra Virginia.[1]

Arten vistas främst i barrskogar men den hittas likaså i blandskogar, buskskogar och urbaniserade områden.[1]

Ekologi

Individerna är aktiva på dagen. På natten vilar de i trädens håligheter, självbyggda bon av blad eller i jordhålor.[1] Vid sämre väderförhållanden blir ekorren slö men den håller ingen vinterdvala. Den samlar vanligen ett stort förråd för dessa tider och ofta används förrådet av flera generationer. Utanför parningstiden lever varje individ ensam och försvarar ett revir kring förrådet.[10]

Amerikansk röd ekorre äter främst frön från kottar samt andra växtdelar som nötter, bark, sav, nektar, blommor och unga växtskott. Denna kost kompletteras med insekter, fågelägg samt mindre ofta med små ryggradsdjur som småfåglar och andra gnagare.[10]

Honorna parar sig vanligen under våren (februari till april) och ibland även under sensommaren (juni – augusti). En hona parar sig oftast med flera hannar. Dräktigheten varar 31 till 35 dagar och sedan föds 3 till 5 ungar. Ungdjuren är vid födelsen nakna. De får di under cirka 7 till 8 veckor och blir självständiga efter ungefär 18 veckor. Ibland lämnar modern i stället för ungarna reviret när ungarna är tillräckligt stora. De flesta individer blir 3 till 7 år gamla och vissa exemplar blir 10 år gamla.[10]

Underarter

Arten delas av Catalogue of Life in i följande 25 underarter:[8]

  • T. h. hudsonicus (Erxleben, 1777)
  • T. h. abieticola (A. H. Howell, 1929)
  • T. h. baileyi (J. A. Allen, 1898)
  • T. h. columbiensis A. H. Howell, 1936
  • T. h. dakotensis (J. A. Allen, 1894)
  • T. h. dixiensis Hardy, 1942
  • T. h. fremonti (Audubon and Bachman, 1853)
  • T. h. grahamensis (J. A. Allen, 1894)
  • T. h. gymnicus (Bangs, 1899)
  • T. h. kenaiensis A. H. Howell, 1936
  • T. h. lanuginosus (Bachman, 1839)
  • T. h. laurentianus Anderson, 1942
  • T. h. loquax (Bangs, 1896)
  • T. h. lychnuchus (Stone and Rehn, 1903)
  • T. h. minnesota (J. A. Allen, 1899)
  • T. h. mogollonensis (Mearns, 1890)
  • T. h. pallescens A. H. Howell, 1942
  • T. h. petulans (Osgood, 1900)
  • T. h. picatus (Swarth, 1921)
  • T. h. preblei A. H. Howell, 1936
  • T. h. regalis A. H. Howell, 1936
  • T. h. richardsoni (Bachman, 1839)
  • T. h. streatori (J. A. Allen, 1898)
  • T. h. ungavensis Anderson, 1942
  • T. h. ventorum (J. A. Allen, 1898)

Wilson & Reeder (2005) räknar inte in T. h. columbiensis.[5]

Källor

  1. ^ [a b c d e] 2008 Tamiasciurus hudsonicus Från: IUCN 2012. IUCN Red List of Threatened Species. Version 2012.2 <www.iucnredlist.org>. Läst 2012-10-24.
  2. ^ (1996) , database, NODC Taxonomic Code
  3. ^ Banks, R. C., R. W. McDiarmid, A. L. Gardner, and W. C. Starnes (2003) , Checklist of Vertebrates of the United States, the U.S. Territories, and Canada
  4. ^ Wilson, Don E., and DeeAnn M. Reeder, eds. (1992) , Mammal Species of the World: A Taxonomic and Geographic Reference, 2nd ed., 3rd printing
  5. ^ [a b] (2005) , website Tamiasciurus hudsonicus, Mammal Species of the World
  6. ^ Wilson, Don E., and F. Russell Cole (2000) , Common Names of Mammals of the World
  7. ^ Banks, R. C., R. W. McDiarmid, and A. L. Gardner (1987) Checklist of Vertebrates of the United States, the U.S. Territories, and Canada, Resource Publication, no. 166
  8. ^ [a b] Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P.M., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L. (red.) (2017). ”Species 2000 & ITIS Catalogue of Life: 2017 Annual Checklist.”. Species 2000: Naturalis, Leiden, Nederländerna. http://www.catalogueoflife.org/annual-checklist/2017/search/all/key/Tamiasciurus+hudsonicus/match/1. Läst 17 november 2017.
  9. ^ ITIS: The Integrated Taxonomic Information System. Orrell T. (custodian), 2011-04-26
  10. ^ [a b c d e] R. Edwards (27 april 2009). ”American red squirrel”. ARKive. Arkiverad från originalet den 16 september 2013. https://web.archive.org/web/20130916030248/http://www.arkive.org/american-red-squirrel/tamiasciurus-hudsonicus/. Läst 19 november 2013.

Externa länkar

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Amerikansk röd ekorre: Brief Summary ( шведски )

добавил wikipedia SV


Amerikansk röd ekorre (Tamiasciurus hudsonicus) är en däggdjursart som först beskrevs av Johann Christian Polycarp Erxleben 1777. Den ingår i släktet Tamiasciurus och familjen ekorrar. IUCN kategoriserar arten globalt som livskraftig.

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Amerika kızıl sincabı ( турски )

добавил wikipedia TR

Amerika kızıl cincabı (Tamiasciurus hudsonicus), sincapgiller familyasından, Kanada ve Amerika Birleşik Devletleri'ninde daha çok Rocky Dağlarında yaşayan, kozalak tohumlarıyla beslenmeye özelleşmiş, gündüzcül bir sincap türü.

Hayvanlarca kullanımı

Amerika kızıl sincapları Kanada vaşağı (Lynx canadensis), doru vaşak (Lynx rufus), kır kurdu (Canis latrans), Amerika puhusu (Bubo virginianus), bayağı çakır kuşu (Accipiter gentilis), kızıl kuyruklu şahin (Buteo jamaicensis), Amerika kargası (Corvus brachyrynchos), Amerika sansarı (Martes americana), kızıl tilki (Vulpes vulpes), boz tilki (Urocyon cinereoargenteus), boz kurt (Canis lupus) ve gelincik (Mustela sp.) trürleri tarafından avlanır.

Alt türleri

  • T. h. hudsonicus
  • T. h. abieticola
  • T. h. baileyi
  • T. h. dakotensis
  • T. h. dixiensis
  • T. h. fremonti
  • T. h. grahamensis
  • T. h. gymnicus
  • T. h. kenaiensis
  • T. h. lanuginosus
  • T. h. laurentianus
  • T. h. loquax
  • T. h. lychnuchus
  • T. h. minnesota
  • T. h. mogollonensis
  • T. h. pallescens
  • T. h. petulans
  • T. h. picatus
  • T. h. preblei
  • T. h. regalis
  • T. h. richardsoni
  • T. h. streatori
  • T. h. ungavensis
  • T. h. ventorum
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Amerika kızıl sincabı: Brief Summary ( турски )

добавил wikipedia TR

Amerika kızıl cincabı (Tamiasciurus hudsonicus), sincapgiller familyasından, Kanada ve Amerika Birleşik Devletleri'ninde daha çok Rocky Dağlarında yaşayan, kozalak tohumlarıyla beslenmeye özelleşmiş, gündüzcül bir sincap türü.

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Tamiasciurus hudsonicus ( украински )

добавил wikipedia UK

Поширення

Країни поширення: Канада (Альберта, Британська Колумбія, Лабрадор, Манітоба, Нью-Брансвік, Північно-Західні території, Нова Шотландія, Нунавут, Онтаріо, Острів Принца Едварда, Квебек, Саскачеван, Юкон), США (Аляска, Аризона, Колорадо, Коннектикут, Делавер, район Колумбія, Джорджія, Айдахо, Іллінойс, Індіана, Айова, Кентуккі, Мен, Меріленд, Массачусетс, Мічиган, Міннесота, Монтана, Нью-Гемпшир, Нью-Джерсі, Нью-Мексико, Нью Йорк, Північна Кароліна, Північна Дакота, Огайо, Орегон, Пенсильванія , Род-Айленд, Південна Кароліна, Південна Дакота, Теннессі, Юта, Вермонт, Вірджинія, Вашингтон, Західна Вірджинія, Вісконсин, Вайомінг). Віддає перевагу хвойним та змішаним лісам, але зустрічається й у листяних лісах.

Життя

Живе поодинці. Віддає перевагу гніздуванню в дуплах дерев, а також будує листяні гнізда і використовує земляні нори. Вагітність триває 31-35 днів. Деякі самиці дають два приплоди на рік, в середньому 4-5 дитинчат у виводку. Раціон складається з насіння, хвойних шишок, горіхів, фруктів. Іноді живиться безхребетними і дрібними хребетними. Зазвичай запасає, а потім споживає, велику кількість їжі. Досить активні протягом дня. Піки денної активності через дві години після сходу сонця і за дві години до заходу.

Морфологічні особивості

Має червонувато-коричневе хутро на спині, з білим черевом. Довжина тіла 17-23 см, довжина хвоста 9-16 см, вуха 2—3 см, вага 197-282 гр. Зубна формула: 1/1, 0/0, 1-2/1, 3/3, загалом 20, дуже рідко 22 зуба. 2n=46.

Джерела


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Tamiasciurus hudsonicus ( виетнамски )

добавил wikipedia VI

Tamiasciurus hudsonicus là một loài động vật có vú trong họ Sóc, bộ Gặm nhấm. Loài này được Erxleben mô tả năm 1777.[2]

Hình ảnh

Chú thích

  1. ^ Linzey, A. V. & Hammerson, G. (2008). Tamiasciurus hudsonicus. 2008 Sách đỏ IUCN. Liên minh Bảo tồn Thiên nhiên Quốc tế 2008. Truy cập ngày 8 tháng 1 năm 2009.
  2. ^ a ă Wilson, D. E.; Reeder, D. M. biên tập (2005). “Tamiasciurus hudsonicus”. Mammal Species of the World . Baltimore: Nhà in Đại học Johns Hopkins, 2 tập (2.142 trang). ISBN 978-0-8018-8221-0. OCLC 62265494.

Tham khảo

  • Thorington, R. W. Jr. and R. S. Hoffman. 2005. Family Sciuridae. pp. 754–818 in Mammal Species of the World a Taxonomic and Geographic Reference. D. E. Wilson and D. M. Reeder eds. Johns Hopkins University Press, Baltimore.
  •  src= Phương tiện liên quan tới Tamiasciurus hudsonicus tại Wikimedia Commons


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Tamiasciurus hudsonicus: Brief Summary ( виетнамски )

добавил wikipedia VI

Tamiasciurus hudsonicus là một loài động vật có vú trong họ Sóc, bộ Gặm nhấm. Loài này được Erxleben mô tả năm 1777.

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Красная белка ( руски )

добавил wikipedia русскую Википедию
Царство: Животные
Подцарство: Эуметазои
Без ранга: Вторичноротые
Подтип: Позвоночные
Инфратип: Челюстноротые
Надкласс: Четвероногие
Подкласс: Звери
Инфракласс: Плацентарные
Надотряд: Euarchontoglires
Грандотряд: Грызунообразные
Отряд: Грызуны
Подотряд: Белкообразные
Семейство: Беличьи
Подсемейство: Sciurinae
Триба: Sciurini
Вид: Красная белка
Международное научное название

Tamiasciurus hudsonicus (Erxleben, 1777)

Подвиды[1]
  • T. h. hudsonicus Erxleben, 1777
  • T. h. abieticola A. H. Howell, 1929
  • T. h. baileyi J. A. Allen, 1898
  • T. h. dacotensis J. A. Allen, 1894
  • T. h. dixiensis Hardy, 1942
  • T. h. fremonti Audubon & Bachman, 1853
  • T. h. grahamensis J. A. Allen, 1894
  • T. h. gymnicus Bangs, 1899
  • T. h. kenaiensis A. H. Howell, 1936
  • T. h. lanuginosus Bachman, 1839
  • T. h. laurentianus Anderson, 1942
  • T. h. loquax Bangs, 1896
  • T. h. lychnuchus Stone & Rehn, 1903
  • T. h. minnesota J. A. Allen, 1899
  • T. h. mogollonensis Mearns, 1890
  • T. h. palescens A. H. Howell, 1942
  • T. h. petulans Osgood, 1900
  • T. h. picatus Swarth, 1921
  • T. h. preblei A. H. Howell, 1936
  • T. h. regalis A. H. Howell, 1936
  • T. h. richardsoni Bachman, 1839
  • T. h. streatori J. A. Allen, 1898
  • T. h. ungavensis Anderson, 1942
  • T. h. ventorum J. A. Allen, 1898
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Систематика
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Изображения
на Викискладе
ITIS 180166NCBI 10009EOL 347433FW 52035

Красная белка[2] (лат. Tamiasciurus hudsonicus) — вид грызунов семейства беличьих, наиболее распространённый и характерный представитель рода красных белок. Научное видовое название «hudsonicus» дано животному в честь Гудзонова залива, места, в котором оно было впервые описано[3].

Распространение и места обитания

Распространён практически по всей территории Северной Америки — на Аляске, в Канаде, в США вплоть до пустынь Аризоны и Нью-Мексико на юге и Джорджии на юго-востоке[4]. Белки населяют различные лесистые местности, в том числе хвойные, лиственные и смешанные леса. Также их можно наблюдать в пригородных районах, где имеются насаждения из больших зрелых деревьев[5].

Описание

Длина тела белок 28—35 см, длина хвоста — 9,5—15 см. Окрас меха очень изменчив[5]. В разных регионах своего ареала эти белки могут иметь вариабельный окрас[5], а также белки меняют мех на зимний и летний[3][5]. Мех обычно коричневатый или оливково-красный. Летом по бокам есть чёрная продольная полоса, которая разделяет живот и спину. Мех на животе белый или кремовый. Хвост обычно имеет белую кайму. Вокруг чёрных глаз мех белый[5].

Экология

В дикой природе красные белки живут до семи лет, но большинство погибают прежде, чем доживут до года. Ведут одиночный и дневной образ жизни; активны круглый год. Активнее всего на рассвете и после полудня. Своё логово размещают в старых дуплах дятлов, древесных пустотах или других мелких расщелинах. На севере ареала красные белки зиму часто проводят в подземной системе тоннелей. Белки часто мигрируют если запас пищи на проживаемой территории снижается. Мигрируя, им часто приходится переплывать водоёмы[5].

В массовой культуре

Красная белка — один из отрицательных персонажей мультсериала Пингвины из Мадагаскара.

Красная белка Чиву (Chewoo) - один из главных персонажей мультсериала "Юху и его друзья" (YooHoo & Friends)

Галерея

  • Gilles Gonthier - Tamiasciurus hudsonicus (by).jpg
  • Gilles Gonthier - Un instant de repos -- A little rest (by).jpg
  • Tamiasciurus hudsonicus 001.jpg
  • Gilles Gonthier - Écureuil roux -- Red Squirrel (by).jpg
  • American Red Squirrel .jpg
  • Swimming red squirrel.jpg
  • Red squirrel with nut.jpg
  • Red squirrel.jpg
  • Tamiasciurus hudsonicus CT.jpg

Примечания

  1. Tamiasciurus hudsonicus (англ.). Wilson & Reeder`s Mammal Species of the World Third Edition. Проверено 16 июля 2011. Архивировано 14 августа 2012 года.
  2. Соколов В. Е. Пятиязычный словарь названий животных. Млекопитающие. Латинский, русский, английский, немецкий, французский. / под общей редакцией акад. В. Е. Соколова. — М.: Рус. яз., 1984. — С. 145. — 10 000 экз.
  3. 1 2 Joseph F. Merritt & Ruth Anne Matinko. Guide to the Mammals of Pennsylvania. — Pittsburgh: University of Pittsburgh Press, 1987. — С. 408. — 152—154 с. — ISBN 0-8229-5393-5.
  4. Tamiasciurus hudsonicus (англ.). The IUCN Red List of Threatened Species.
  5. 1 2 3 4 5 6 Dewey, T. & E. Ellis. Информация о виде (англ.). University of Michigan Museum of Zoology: Animal Diversity Web (animaldiversity.ummz.umich.edu). Проверено 17 июля 2011. Архивировано 14 августа 2012 года.


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Красная белка: Brief Summary ( руски )

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Красная белка (лат. Tamiasciurus hudsonicus) — вид грызунов семейства беличьих, наиболее распространённый и характерный представитель рода красных белок. Научное видовое название «hudsonicus» дано животному в честь Гудзонова залива, места, в котором оно было впервые описано.

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北美紅松鼠 ( кинески )

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北美紅松鼠: Brief Summary ( кинески )

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北美紅松鼠(學名:Tamiasciurus hudsonicus)是美洲紅松鼠的其中一種。

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아메리카붉은다람쥐청서 ( корејски )

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아메리카붉은다람쥐 또는 아메리카붉은청서(영어: American red squirrel, Tamiasciurus hudsonicus)는 다람쥐과에 속하는 설치류의 일종이다.[2] 타미아스키우루스속에 속하는 3종의 나무 다람쥐의 하나로 나머지 종은 더글라스다람쥐청서먼스다람쥐청서이다. 몸무게 200~250g의 중형 크기의 청서류로 낮에 활동하는 주행성 포유류이다. 일년 내내 독점적인 영역에서 활동을 한다. 천적은 여우, 캐나다스라소니, 대형맹금류, 이다.

아종

24종의 아종이 알려져 있다.[2]

각주

  1. Cassola, F. (2016). Tamiasciurus hudsonicus. The IUCN Red List of Threatened Species doi 10.2305/IUCN.UK.2016-3.RLTS.T42587A22250817.en
  2. Thorington, R.W., Jr.; Hoffman, R.S. (2005). 〈Family Sciuridae〉 [다람쥐과]. Wilson, D.E.; Reeder, D.M. 《Mammal Species of the World: A Taxonomic and Geographic Reference》 (영어) 3판. 존스 홉킨스 대학교 출판사. 754–818쪽. ISBN 978-0-8018-8221-0. OCLC 62265494.
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아메리카붉은다람쥐청서: Brief Summary ( корејски )

добавил wikipedia 한국어 위키백과

아메리카붉은다람쥐 또는 아메리카붉은청서(영어: American red squirrel, Tamiasciurus hudsonicus)는 다람쥐과에 속하는 설치류의 일종이다. 타미아스키우루스속에 속하는 3종의 나무 다람쥐의 하나로 나머지 종은 더글라스다람쥐청서먼스다람쥐청서이다. 몸무게 200~250g의 중형 크기의 청서류로 낮에 활동하는 주행성 포유류이다. 일년 내내 독점적인 영역에서 활동을 한다. 천적은 여우, 캐나다스라소니, 대형맹금류, 이다.

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